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mimule musqué, musk monkeyflower, musk-flower, musk-plant, sticky monkey-flower

Utah monkeyflower

Habit Perennials, rhizomatous, rooting at proximal nodes. Perennials, rhizomatous, rooting at nodes, rhizomes filiform.
Stems

erect, sometimes ascending to decumbent, simple or branched, (2–)5–20 cm, nodes 2–4(or 5), glabrate to glandular-villous, hairs 0.5–2 mm, gland-tipped, internodes evident.

erect, sometimes decumbent-ascending proximally, simple or few-branched, 20–50 cm, glabrous or sparsely stipitate-glandular in inflorescence.

Leaves

usually cauline, basal not persistent, distinctly separated;

petiole 0 mm or (0.5–)1–5(–10) mm;

blade pinnately veined, oblong-ovate to ovate, (10–)15–40(–50) × 5–25 mm, base obtuse-cuneate to truncate, rounded or subcordate, subclasping to sessile, margins coarsely serrate-dentate to denticulate or subentire, apex acute to obtuse, surfaces glabrate to glandular-villous.

basal and cauline, even-sized or largest near mid stem;

petiole 0 mm or proximalmost 2–10 mm;

blade palmately 3–5-veined, orbicular or suborbicular to broadly elliptic, broadly ovate, or depressed-ovate, 20–40(–75) × 12–35(–40) mm, base usually truncate to broadly cuneate, margins entire or subentire to mucronulate, shallowly dentate, or denticulate, apex rounded, surfaces glabrous or glabrate to sparsely stipitate-glandular and sparsely pilose, hairs thin-walled, abaxial often glaucous.

Flowers

herkogamous, 1–8, from medial to distal nodes.

herkogamous, 6–16, from proximal to distal nodes, in a loose raceme, distal bracts becoming much reduced.

Styles

glabrous.

hirtellous.

Corollas

yellow, throat with fine red to blackish or brown lines extending onto lobes, red to brown dots in throat and lobes present or absent, bilaterally or nearly radially symmetric, bilabiate or nearly regular;

tube-throat narrowly funnelform, 11–18 mm, exserted beyond calyx margin;

lobes oblong-obovate, apex rounded to notched.

yellow, abaxial limb prominently darker yellow, sparsely purple-spotted, bilaterally symmetric, weakly bilabiate;

tube-throat narrowly funnelform to broadly cylindric, 10–15 mm, exserted 5–8 mm beyond calyx margin;

limb expanded 12–20 mm.

Fruiting pedicels

(7–)10–25 mm, glabrate to glandular-villous.

(25–)40–75 mm, stipitate-glandular to short villous, hairs gland-tipped.

Fruiting calyces

ridge- to wing-angled, campanulate to cylindric-campanulate, weakly or not inflated, 6–13 mm, villous to glandular-villous, lobes erect to spreading-recurving, strongly unequal to subequal, triangular to linear-lanceolate or narrowly triangular-acuminate, 2–4 mm, apex acute to obtuse.

broadly ovate-cylindric, inflated, sagittally compressed, (10–)11–17(–20) mm, stipitate-glandular or minutely hirtellous or a mixture, hairs sometimes also longer, thin-walled, eglandular or glandular, throat not closing, adaxial lobe slightly longer, triangular-blunt.

Capsules

included, 6–8 mm.

included, 5–7 mm.

Anthers

included, glabrous or slightly hirtellous to scabrous.

included, glabrous.

2n

= 32.

= 28, 30.

Erythranthe moschata

Erythranthe utahensis

Phenology Flowering May–Aug. Flowering (Feb–)May–Aug(–Oct).
Habitat Springs and seeps, creek edges, moist meadows, ditches, along trails, roadsides, rocky ridges, granite outcrops, shaded and wet places in sagebrush, aspen, fir, spruce-fir, lodgepole pine forests, meadows. Drainage ditches, springs, seeps, wet meadows, margins of ponds and small streams, marshy areas.
Elevation (300–)400–3100 m. ((1000–)1300–10200 ft.) 1400–2500 m. (4600–8200 ft.)
Distribution
from FNA
CA; CO; CT; ID; MA; ME; MI; MT; NH; NJ; NV; NY; OR; PA; RI; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM [Introduced in South America (Chile), Europe, e Asia (Japan), Pacific Islands (New Zealand), Australia]
[WildflowerSearch map]
from FNA
CA; CO; NV; UT
Discussion

Earlier segregation of Erythranthe moniliformis as distinct from E. moschata (for example, G. L. Nesom 2012g) emphasized a primarily erect habit and tendency toward sessile to subsessile and more densely arranged cauline leaves in E. moniliformis versus a decumbent to procumbent habit and consistently petiolate leaves on longer internodes in E. moschata. Discontinuities in morphology, geography, and ecology were not confirmed in later study by Nesom (2017). Rhizomes with small, tuberlike swellings can be observed over the whole moschata/moniliformis range, and there apparently are no consistent distinctions in vestiture and corolla size.

Mimulus acutidens Reiche (1911), a later homonym of M. acutidens Greene, pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe utahensis is characterized by its erect stems, prolifically produced filiform rhizomes, basal leaves short-petiolate to subsessile and cauline sessile, blades suborbicular to broadly ovate or depressed ovate with thin-walled villous-glandular hairs on both surfaces, margins subentire, proximal pedicels elongating to 75 mm, and calyces open at maturity.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 401. FNA vol. 17, p. 410.
Parent taxa Phrymaceae > Erythranthe Phrymaceae > Erythranthe
Sibling taxa
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. verbenacea, E. washingtonensis, E. willisii
Synonyms Mimulus moschatus, E. inodora, E. moniliformis, M. crinitus, M. guttatus var. moschatus, M. inodorus, M. leibergii, M. macranthus, M. moniliformis, M. moschatus var. longiflorus, M. moschatus var. moniliformis, M. moschatus var. pallidiflorus Mimulus glabratus subsp. utahensis
Name authority (Douglas ex Lindley) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) (Pennell) G. L. Nesom: Phytoneuron 2012-39: 44. (2012)
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