Erythranthe moschata |
Erythranthe regni |
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mimule musqué, musk monkeyflower, musk-flower, musk-plant, sticky monkey-flower |
King of Arizona monkeyflower |
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Habit | Perennials, rhizomatous, rooting at proximal nodes. | Annuals, fibrous-rooted, sometimes rooting at proximal nodes. |
Stems | erect, sometimes ascending to decumbent, simple or branched, (2–)5–20 cm, nodes 2–4(or 5), glabrate to glandular-villous, hairs 0.5–2 mm, gland-tipped, internodes evident. |
erect to ascending-erect, branched, sometimes becoming slightly fistulose, 15–45 cm, glabrous. |
Leaves | usually cauline, basal not persistent, distinctly separated; petiole 0 mm or (0.5–)1–5(–10) mm; blade pinnately veined, oblong-ovate to ovate, (10–)15–40(–50) × 5–25 mm, base obtuse-cuneate to truncate, rounded or subcordate, subclasping to sessile, margins coarsely serrate-dentate to denticulate or subentire, apex acute to obtuse, surfaces glabrate to glandular-villous. |
basal and cauline; petiole: proximals 5–25(–30) mm, mid cauline and distals not connate, 0 mm; blade palmately 5–7-veined, proximal sometimes subpinnate, proximals ovate to depressed-orbicular, 15–20(–50) × 15–25(–50) mm, medials and distals broadly depressed-ovate to obtriangular or flabellate, 15–35 mm, largest basal or at mid stem with distal slightly reduced, base attenuate-cuneate, margins shallowly serrate-dentate, sometimes irregularly, to mucronulate or apiculate, teeth (3–)5–7 per side, rarely subentire, apex rounded, surfaces glabrous. |
Flowers | herkogamous, 1–8, from medial to distal nodes. |
plesiogamous, 6–16, from all nodes or medial to distal, cleistogamous. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine red to blackish or brown lines extending onto lobes, red to brown dots in throat and lobes present or absent, bilaterally or nearly radially symmetric, bilabiate or nearly regular; tube-throat narrowly funnelform, 11–18 mm, exserted beyond calyx margin; lobes oblong-obovate, apex rounded to notched. |
yellow, not red-dotted, bilaterally or radially symmetric, bilabiate or regular; tube-throat cylindric-funnelform, 9–12 mm, exserted 3–5 mm beyond calyx margin; limb expanded 1–1.5 mm. |
Fruiting pedicels | (7–)10–25 mm, glabrate to glandular-villous. |
15–30 mm, longer than subtending leaves, glabrous. |
Fruiting calyces | ridge- to wing-angled, campanulate to cylindric-campanulate, weakly or not inflated, 6–13 mm, villous to glandular-villous, lobes erect to spreading-recurving, strongly unequal to subequal, triangular to linear-lanceolate or narrowly triangular-acuminate, 2–4 mm, apex acute to obtuse. |
sparsely purple-dotted, broadly campanulate-cylindric, inflated, sagittally compressed, 7–9 mm, glabrous, throat not closing, adaxial lobe longest. |
Capsules | included, 6–8 mm. |
included, 4–5 mm. |
Anthers | included, glabrous or slightly hirtellous to scabrous. |
included, glabrous. |
2n | = 32. |
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Erythranthe moschata |
Erythranthe regni |
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Phenology | Flowering May–Aug. | Flowering Mar–May. |
Habitat | Springs and seeps, creek edges, moist meadows, ditches, along trails, roadsides, rocky ridges, granite outcrops, shaded and wet places in sagebrush, aspen, fir, spruce-fir, lodgepole pine forests, meadows. | Moist to wet, sandy loam soils. |
Elevation | (300–)400–3100 m. ((1000–)1300–10200 ft.) | 800–1000 m. (2600–3300 ft.) |
Distribution |
CA; CO; CT; ID; MA; ME; MI; MT; NH; NJ; NV; NY; OR; PA; RI; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM [Introduced in South America (Chile), Europe, e Asia (Japan), Pacific Islands (New Zealand), Australia]
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AZ |
Discussion | Earlier segregation of Erythranthe moniliformis as distinct from E. moschata (for example, G. L. Nesom 2012g) emphasized a primarily erect habit and tendency toward sessile to subsessile and more densely arranged cauline leaves in E. moniliformis versus a decumbent to procumbent habit and consistently petiolate leaves on longer internodes in E. moschata. Discontinuities in morphology, geography, and ecology were not confirmed in later study by Nesom (2017). Rhizomes with small, tuberlike swellings can be observed over the whole moschata/moniliformis range, and there apparently are no consistent distinctions in vestiture and corolla size. Mimulus acutidens Reiche (1911), a later homonym of M. acutidens Greene, pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe regni is endemic to the Kofa Mountains of Yuma County; all collections have been made from the Kofa Game Refuge (Kofa National Wildlife Refuge). Because its calyces remain open at maturity, this species is hypothesized to be most closely related to E. geyeri, from which it differs by its erect habit, apparently annual duration, larger leaves, purple-dotted calyces, and corollas with longer tube-throat and barely bilabiate limb. Geography and other morphology, however, suggest that its evolutionary origins are closer to E. guttata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 401. | FNA vol. 17, p. 406. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus, E. inodora, E. moniliformis, M. crinitus, M. guttatus var. moschatus, M. inodorus, M. leibergii, M. macranthus, M. moniliformis, M. moschatus var. longiflorus, M. moschatus var. moniliformis, M. moschatus var. pallidiflorus | |
Name authority | (Douglas ex Lindley) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | G. L. Nesom: Phytoneuron 2012-40: 24. (2012) |
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