Erythranthe moschata |
Erythranthe laciniata |
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mimule musqué, musk monkeyflower, musk-flower, musk-plant, sticky monkey-flower |
cut-leaf monkeyflower |
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Habit | Perennials, rhizomatous, rooting at proximal nodes. | Annuals, slender-taprooted or fibrous-rooted. |
Stems | erect, sometimes ascending to decumbent, simple or branched, (2–)5–20 cm, nodes 2–4(or 5), glabrate to glandular-villous, hairs 0.5–2 mm, gland-tipped, internodes evident. |
erect, simple or branched from base, 3–38 cm, glabrous or sparsely hirtellous, finely villosulous-glandular above nodes. |
Leaves | usually cauline, basal not persistent, distinctly separated; petiole 0 mm or (0.5–)1–5(–10) mm; blade pinnately veined, oblong-ovate to ovate, (10–)15–40(–50) × 5–25 mm, base obtuse-cuneate to truncate, rounded or subcordate, subclasping to sessile, margins coarsely serrate-dentate to denticulate or subentire, apex acute to obtuse, surfaces glabrate to glandular-villous. |
cauline, basal deciduous by flowering; petiole 1–35 mm, distals 0 mm; blade 1-veined or palmately 3-veined, elliptic to elliptic-obovate, oblanceolate, or oblong, 3–55 mm, longer than wide, base attenuate, margins narrowly pinnately lobed or dissected, sometimes merely shallowly toothed, apex acute to obtuse, surfaces glabrate. |
Flowers | herkogamous, 1–8, from medial to distal nodes. |
plesiogamous, 2–8, from medial to distal nodes, chasmogamous, sometimes cleistogamous. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine red to blackish or brown lines extending onto lobes, red to brown dots in throat and lobes present or absent, bilaterally or nearly radially symmetric, bilabiate or nearly regular; tube-throat narrowly funnelform, 11–18 mm, exserted beyond calyx margin; lobes oblong-obovate, apex rounded to notched. |
yellow, throat red-spotted, abaxial limb of larger usually with 1 large red splotch, bilaterally symmetric, ± bilabiate; tube-throat funnelform, 4–6 mm, exserted 1–2 mm beyond calyx margin; limb expanded 5–6 mm. |
Fruiting pedicels | (7–)10–25 mm, glabrate to glandular-villous. |
nodding 30–140º at calyx base, 5–25 mm. |
Fruiting calyces | ridge- to wing-angled, campanulate to cylindric-campanulate, weakly or not inflated, 6–13 mm, villous to glandular-villous, lobes erect to spreading-recurving, strongly unequal to subequal, triangular to linear-lanceolate or narrowly triangular-acuminate, 2–4 mm, apex acute to obtuse. |
red-spotted, cylindric-campanulate, inflated, sagittally compressed, 8–10 mm, glabrate, throat closing, lobes ca. equal size or adaxial slightly longer. |
Capsules | included, 6–8 mm. |
included, stipitate, 5–7 mm. |
Anthers | included, glabrous or slightly hirtellous to scabrous. |
included, glabrous. |
2n | = 32. |
= 28. |
Erythranthe moschata |
Erythranthe laciniata |
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Phenology | Flowering May–Aug. | Flowering Apr–Jul(–Aug). |
Habitat | Springs and seeps, creek edges, moist meadows, ditches, along trails, roadsides, rocky ridges, granite outcrops, shaded and wet places in sagebrush, aspen, fir, spruce-fir, lodgepole pine forests, meadows. | Cracks, depressions, and seeps in granite outcrops, ledges, talus and scree, rocky streamsides, rocky slopes, roadsides, intermittent drainages. |
Elevation | (300–)400–3100 m. ((1000–)1300–10200 ft.) | 900–2300(–3300) m. (3000–7500(–10800) ft.) |
Distribution |
CA; CO; CT; ID; MA; ME; MI; MT; NH; NJ; NV; NY; OR; PA; RI; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM [Introduced in South America (Chile), Europe, e Asia (Japan), Pacific Islands (New Zealand), Australia]
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CA
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Discussion | Earlier segregation of Erythranthe moniliformis as distinct from E. moschata (for example, G. L. Nesom 2012g) emphasized a primarily erect habit and tendency toward sessile to subsessile and more densely arranged cauline leaves in E. moniliformis versus a decumbent to procumbent habit and consistently petiolate leaves on longer internodes in E. moschata. Discontinuities in morphology, geography, and ecology were not confirmed in later study by Nesom (2017). Rhizomes with small, tuberlike swellings can be observed over the whole moschata/moniliformis range, and there apparently are no consistent distinctions in vestiture and corolla size. Mimulus acutidens Reiche (1911), a later homonym of M. acutidens Greene, pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe laciniata is known from Amador County south to Kern County. As in Erythranthe nasuta, the adaxial calyx lobe in E. laciniata tends to be narrowly lanceolate to triangular (noselike) and perceptibly falcate, curving slightly upward both in flower and in fruit. The adaxial lobe is not so prominently protruding as it often is in E. nasuta. Corolla size is variable in Erythranthe laciniata, but the size of those with an open throat (versus much reduced in size and apparently cleistogamous) is not strongly correlated with size of the individual plant, and all on one plant are about the same size (compare with E. nasuta). Corollas on some plants, however, are all or nearly all greatly reduced and apparently cleistogamous. Fertilization in even the larger corollas apparently is autogamous; the anther pairs are slightly separated or equal in level, and the stigma is in the middle of the anthers or at the level of the adaxial pair. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 401. | FNA vol. 17, p. 419. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus, E. inodora, E. moniliformis, M. crinitus, M. guttatus var. moschatus, M. inodorus, M. leibergii, M. macranthus, M. moniliformis, M. moschatus var. longiflorus, M. moschatus var. moniliformis, M. moschatus var. pallidiflorus | Mimulus laciniatus, M. eisenii |
Name authority | (Douglas ex Lindley) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | (A. Gray) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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