Erythranthe moschata |
Erythranthe inflatula |
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mimule musqué, musk monkeyflower, musk-flower, musk-plant, sticky monkey-flower |
disappearing monkey-flower, ephemeral monkeyflower |
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Habit | Perennials, rhizomatous, rooting at proximal nodes. | Annuals, fibrous-rooted or filiform-taprooted. |
Stems | erect, sometimes ascending to decumbent, simple or branched, (2–)5–20 cm, nodes 2–4(or 5), glabrate to glandular-villous, hairs 0.5–2 mm, gland-tipped, internodes evident. |
erect to ascending, straight or geniculate at nodes, simple or branched at proximal and medial nodes, 6–20(–25) cm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
Leaves | usually cauline, basal not persistent, distinctly separated; petiole 0 mm or (0.5–)1–5(–10) mm; blade pinnately veined, oblong-ovate to ovate, (10–)15–40(–50) × 5–25 mm, base obtuse-cuneate to truncate, rounded or subcordate, subclasping to sessile, margins coarsely serrate-dentate to denticulate or subentire, apex acute to obtuse, surfaces glabrate to glandular-villous. |
usually cauline, basal usually deciduous by flowering; petiole: proximals 1–3 mm, distals 0 mm; blade palmately 3–5-veined, narrowly ovate or narrowly lanceolate to elliptic or elliptic-lanceolate, largest 8–18(–30) × (1–)3–7 mm, relatively even-sized, or slightly reduced distally, base attenuate to obtuse or rounded, margins entire, mucronulate, or denticulate, apex acute to obtuse, surfaces minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
Flowers | herkogamous, 1–8, from medial to distal nodes. |
plesiogamous, 10–20, from medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine red to blackish or brown lines extending onto lobes, red to brown dots in throat and lobes present or absent, bilaterally or nearly radially symmetric, bilabiate or nearly regular; tube-throat narrowly funnelform, 11–18 mm, exserted beyond calyx margin; lobes oblong-obovate, apex rounded to notched. |
yellow to pale yellow, sparsely red-spotted or not, bilaterally symmetric, weakly bilabiate; tube-throat cylindric, 5–8 mm, exserted 1–3 mm beyond calyx margin; limb barely widened, lobes broadly obovate, apex rounded or mucronate. |
Fruiting pedicels | (7–)10–25 mm, glabrate to glandular-villous. |
straight, 7–18 mm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
Fruiting calyces | ridge- to wing-angled, campanulate to cylindric-campanulate, weakly or not inflated, 6–13 mm, villous to glandular-villous, lobes erect to spreading-recurving, strongly unequal to subequal, triangular to linear-lanceolate or narrowly triangular-acuminate, 2–4 mm, apex acute to obtuse. |
winged, plicate-angled, maturing ovoid-ellipsoid to campanulate or broadly urceolate, distinctly inflated, 7–11 mm, margins distinctly toothed or lobed, sparsely, minutely hirtellous, eglandular, lobes pronounced, erect. |
Capsules | included, 6–8 mm. |
included, 5–9 mm. |
Anthers | included, glabrous or slightly hirtellous to scabrous. |
included, glabrous. |
2n | = 32. |
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Erythranthe moschata |
Erythranthe inflatula |
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Phenology | Flowering May–Aug. | Flowering Jun–Jul. |
Habitat | Springs and seeps, creek edges, moist meadows, ditches, along trails, roadsides, rocky ridges, granite outcrops, shaded and wet places in sagebrush, aspen, fir, spruce-fir, lodgepole pine forests, meadows. | Drying edges, banks, and beds of summer-dry watercourses, near drying edges of small lakes or impoundments, often among rocks and shoreline detritus, occasionally in moist protected areas beneath low shrubs. |
Elevation | (300–)400–3100 m. ((1000–)1300–10200 ft.) | 1200–1700 m. (3900–5600 ft.) |
Distribution |
CA; CO; CT; ID; MA; ME; MI; MT; NH; NJ; NV; NY; OR; PA; RI; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM [Introduced in South America (Chile), Europe, e Asia (Japan), Pacific Islands (New Zealand), Australia]
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CA; ID; NV; OR |
Discussion | Earlier segregation of Erythranthe moniliformis as distinct from E. moschata (for example, G. L. Nesom 2012g) emphasized a primarily erect habit and tendency toward sessile to subsessile and more densely arranged cauline leaves in E. moniliformis versus a decumbent to procumbent habit and consistently petiolate leaves on longer internodes in E. moschata. Discontinuities in morphology, geography, and ecology were not confirmed in later study by Nesom (2017). Rhizomes with small, tuberlike swellings can be observed over the whole moschata/moniliformis range, and there apparently are no consistent distinctions in vestiture and corolla size. Mimulus acutidens Reiche (1911), a later homonym of M. acutidens Greene, pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
No natural occurrences of Erythranthe inflatula are known from Washington; the type collection from Klickitat County is from a cultivated plant. Morphological and molecular data (R. J. Meinke 1995; P. M. Beardsley et al. 2004) indicate that Erythranthe inflatula originated as a hybrid between E. breviflora and E. latidens. Its geography and biology suggest that it is reproductively stable. The putative parents are geographically and ecologically separated for most of their ranges, and the range of E. inflatula is considerably broader than the relatively small region where the parents are sympatric. In the region of sympatry, however, E. inflatula may be difficult to distinguish from one or both of its putative parents. G. L. Nesom (2012g) was not able to find morphology that would distinguish the recently described Mimulus evanescens from E. inflatula. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 401. | FNA vol. 17, p. 400. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus, E. inodora, E. moniliformis, M. crinitus, M. guttatus var. moschatus, M. inodorus, M. leibergii, M. macranthus, M. moniliformis, M. moschatus var. longiflorus, M. moschatus var. moniliformis, M. moschatus var. pallidiflorus | Mimulus inflatulus, M. evanescens |
Name authority | (Douglas ex Lindley) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | (Suksdorf) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) |
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