Erythranthe moschata |
Erythranthe hardhamiae |
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mimule musqué, musk monkeyflower, musk-flower, musk-plant, sticky monkey-flower |
Hardham's monkeyflower, Santa Lucia monkeyflower |
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Habit | Perennials, rhizomatous, rooting at proximal nodes. | Annuals, taprooted. |
Stems | erect, sometimes ascending to decumbent, simple or branched, (2–)5–20 cm, nodes 2–4(or 5), glabrate to glandular-villous, hairs 0.5–2 mm, gland-tipped, internodes evident. |
erect, simple or branched from basal nodes, (2–)3–13 cm, glabrous or minutely puberulent. |
Leaves | usually cauline, basal not persistent, distinctly separated; petiole 0 mm or (0.5–)1–5(–10) mm; blade pinnately veined, oblong-ovate to ovate, (10–)15–40(–50) × 5–25 mm, base obtuse-cuneate to truncate, rounded or subcordate, subclasping to sessile, margins coarsely serrate-dentate to denticulate or subentire, apex acute to obtuse, surfaces glabrate to glandular-villous. |
cauline, basal not persistent; petiole 0 mm; blade palmately 3-veined (in broader ones), linear to oblanceolate, 2–12 × 1–5 mm, base attenuate, margins entire, sometimes toothed, apex acute, surfaces glabrous or minutely puberulent. |
Flowers | herkogamous, 1–8, from medial to distal nodes. |
herkogamous, 1–12, from distal or medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine red to blackish or brown lines extending onto lobes, red to brown dots in throat and lobes present or absent, bilaterally or nearly radially symmetric, bilabiate or nearly regular; tube-throat narrowly funnelform, 11–18 mm, exserted beyond calyx margin; lobes oblong-obovate, apex rounded to notched. |
pink to purple, abaxial limb with 2 yellow palate ridges, bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 5–10 mm, exserted 2–5 mm beyond calyx margin; limb expanded 7–13 mm, lobes notched, abaxial limb sparsely bearded. |
Fruiting pedicels | (7–)10–25 mm, glabrate to glandular-villous. |
ascending to often spreading horizontally, 10–60 mm. |
Fruiting calyces | ridge- to wing-angled, campanulate to cylindric-campanulate, weakly or not inflated, 6–13 mm, villous to glandular-villous, lobes erect to spreading-recurving, strongly unequal to subequal, triangular to linear-lanceolate or narrowly triangular-acuminate, 2–4 mm, apex acute to obtuse. |
becoming reddish, sometimes red-spotted, campanulate to cylindric, 4–6 mm, margins distinctly toothed or lobed, glabrous or minutely puberulent, ribs weak, lobes pronounced, erect, margins glabrous. |
Capsules | included, 6–8 mm. |
included, 4–5 mm. |
Anthers | included, glabrous or slightly hirtellous to scabrous. |
included or slightly exserted, glabrous. |
Stigmas | equal in length to corolla tube or exserted. |
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2n | = 32. |
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Erythranthe moschata |
Erythranthe hardhamiae |
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Phenology | Flowering May–Aug. | Flowering Mar–Apr. |
Habitat | Springs and seeps, creek edges, moist meadows, ditches, along trails, roadsides, rocky ridges, granite outcrops, shaded and wet places in sagebrush, aspen, fir, spruce-fir, lodgepole pine forests, meadows. | Sandy soils near sandstone outcrops and chaparral. |
Elevation | (300–)400–3100 m. ((1000–)1300–10200 ft.) | 300–800 m. (1000–2600 ft.) |
Distribution |
CA; CO; CT; ID; MA; ME; MI; MT; NH; NJ; NV; NY; OR; PA; RI; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM [Introduced in South America (Chile), Europe, e Asia (Japan), Pacific Islands (New Zealand), Australia]
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CA |
Discussion | Earlier segregation of Erythranthe moniliformis as distinct from E. moschata (for example, G. L. Nesom 2012g) emphasized a primarily erect habit and tendency toward sessile to subsessile and more densely arranged cauline leaves in E. moniliformis versus a decumbent to procumbent habit and consistently petiolate leaves on longer internodes in E. moschata. Discontinuities in morphology, geography, and ecology were not confirmed in later study by Nesom (2017). Rhizomes with small, tuberlike swellings can be observed over the whole moschata/moniliformis range, and there apparently are no consistent distinctions in vestiture and corolla size. Mimulus acutidens Reiche (1911), a later homonym of M. acutidens Greene, pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe hardhamiae is endemic to the central coast region in Monterey and San Luis Obispo counties. The species was previously included in E. palmeri but can be distinguished by having a wider limb (16–25 mm) than E. palmeri (8–15 mm) and pale pink flowers with a broad yellow palate and orifice. In contrast, E. palmeri has deep pink flowers with two yellow ridges on the palate. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 401. | FNA vol. 17, p. 388. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus, E. inodora, E. moniliformis, M. crinitus, M. guttatus var. moschatus, M. inodorus, M. leibergii, M. macranthus, M. moniliformis, M. moschatus var. longiflorus, M. moschatus var. moniliformis, M. moschatus var. pallidiflorus | |
Name authority | (Douglas ex Lindley) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | N. S. Fraga: Aliso 30: 64, figs. 23–25. (2012) |
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