Erythranthe moschata |
Erythranthe grayi |
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mimule musqué, musk monkeyflower, musk-flower, musk-plant, sticky monkey-flower |
Gray's monkeyflower, mariposa monkeyflower |
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Habit | Perennials, rhizomatous, rooting at proximal nodes. | Annuals, fibrous-rooted. |
Stems | erect, sometimes ascending to decumbent, simple or branched, (2–)5–20 cm, nodes 2–4(or 5), glabrate to glandular-villous, hairs 0.5–2 mm, gland-tipped, internodes evident. |
erect, simple or branched from base, weakly 4-angled, 8–20 cm, glabrous. |
Leaves | usually cauline, basal not persistent, distinctly separated; petiole 0 mm or (0.5–)1–5(–10) mm; blade pinnately veined, oblong-ovate to ovate, (10–)15–40(–50) × 5–25 mm, base obtuse-cuneate to truncate, rounded or subcordate, subclasping to sessile, margins coarsely serrate-dentate to denticulate or subentire, apex acute to obtuse, surfaces glabrate to glandular-villous. |
basal and cauline; petiole 0 mm; blade palmately 3–5-veined, broadly ovate, 7–18 × 5–12 mm, base rounded, margins denticulate, apex acute, surfaces glabrous, rarely sparsely puberulent. |
Flowers | herkogamous, 1–8, from medial to distal nodes. |
herkogamous, 2–20, from proximal to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine red to blackish or brown lines extending onto lobes, red to brown dots in throat and lobes present or absent, bilaterally or nearly radially symmetric, bilabiate or nearly regular; tube-throat narrowly funnelform, 11–18 mm, exserted beyond calyx margin; lobes oblong-obovate, apex rounded to notched. |
rose red, throat pink lined with rose red and a yellow patch, abaxial ridges yellow, bilaterally symmetric, weakly bilabiate; tube-throat cylindric-funnelform, 8–11 mm, exserted beyond calyx margin; limb expanded 7–10 mm. |
Fruiting pedicels | (7–)10–25 mm, glabrate to glandular-villous. |
6–7 mm, shorter than subtending leaves, glabrous. |
Fruiting calyces | ridge- to wing-angled, campanulate to cylindric-campanulate, weakly or not inflated, 6–13 mm, villous to glandular-villous, lobes erect to spreading-recurving, strongly unequal to subequal, triangular to linear-lanceolate or narrowly triangular-acuminate, 2–4 mm, apex acute to obtuse. |
campanulate, 9–11 × 5–6 mm, margins subtruncate, glabrous, sometimes densely papillate at flowering with tiny, 1-celled, eglandular hairs, these apparently deciduous by fruiting, lobes reduced, subequal. |
Capsules | included, 6–8 mm. |
included, 5–9 mm. |
Anthers | included, glabrous or slightly hirtellous to scabrous. |
included, minutely villous-hirsute. |
2n | = 32. |
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Erythranthe moschata |
Erythranthe grayi |
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Phenology | Flowering May–Aug. | Flowering May–Jul(–Oct). |
Habitat | Springs and seeps, creek edges, moist meadows, ditches, along trails, roadsides, rocky ridges, granite outcrops, shaded and wet places in sagebrush, aspen, fir, spruce-fir, lodgepole pine forests, meadows. | Drying pond beds, creek banks, yellow pine, yellow pine-Libocedrus woodlands. |
Elevation | (300–)400–3100 m. ((1000–)1300–10200 ft.) | 1000–1900 m. (3300–6200 ft.) |
Distribution |
CA; CO; CT; ID; MA; ME; MI; MT; NH; NJ; NV; NY; OR; PA; RI; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM [Introduced in South America (Chile), Europe, e Asia (Japan), Pacific Islands (New Zealand), Australia]
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CA
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Discussion | Earlier segregation of Erythranthe moniliformis as distinct from E. moschata (for example, G. L. Nesom 2012g) emphasized a primarily erect habit and tendency toward sessile to subsessile and more densely arranged cauline leaves in E. moniliformis versus a decumbent to procumbent habit and consistently petiolate leaves on longer internodes in E. moschata. Discontinuities in morphology, geography, and ecology were not confirmed in later study by Nesom (2017). Rhizomes with small, tuberlike swellings can be observed over the whole moschata/moniliformis range, and there apparently are no consistent distinctions in vestiture and corolla size. Mimulus acutidens Reiche (1911), a later homonym of M. acutidens Greene, pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In addition to the features noted in the key and descriptions, the fruiting calyces of Erythranthe grayi are distinctly more inflated than those of E. acutidens and E. inconspicua. G. L. Nesom (2012g) maintained E. grayi as distinct from E. acutidens, relying primarily on fruiting pedicel length (see key above), but the two have nearly identical ranges (Tuolumne County south to Kern County), and study of additional collections suggests that only a single species may be represented. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 401. | FNA vol. 17, p. 382. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus, E. inodora, E. moniliformis, M. crinitus, M. guttatus var. moschatus, M. inodorus, M. leibergii, M. macranthus, M. moniliformis, M. moschatus var. longiflorus, M. moschatus var. moniliformis, M. moschatus var. pallidiflorus | Mimulus grayi |
Name authority | (Douglas ex Lindley) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | (A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 34. (2012) |
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