Erythranthe moschata |
Erythranthe corallina |
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mimule musqué, musk monkeyflower, musk-flower, musk-plant, sticky monkey-flower |
coralline monkeyflower |
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Habit | Perennials, rhizomatous, rooting at proximal nodes. | Perennials, rhizomatous, rhizomes often forming a mass, usually branching, filiform. |
Stems | erect, sometimes ascending to decumbent, simple or branched, (2–)5–20 cm, nodes 2–4(or 5), glabrate to glandular-villous, hairs 0.5–2 mm, gland-tipped, internodes evident. |
usually erect to ascending-erect, few-branched, 6–25(–38) cm, moderately hirsute to hirtellous, hairs deflexed. |
Leaves | usually cauline, basal not persistent, distinctly separated; petiole 0 mm or (0.5–)1–5(–10) mm; blade pinnately veined, oblong-ovate to ovate, (10–)15–40(–50) × 5–25 mm, base obtuse-cuneate to truncate, rounded or subcordate, subclasping to sessile, margins coarsely serrate-dentate to denticulate or subentire, apex acute to obtuse, surfaces glabrate to glandular-villous. |
basal and cauline, becoming larger distally or even-sized; petiole 0 mm or proximals 1–15 mm; blade palmately 5-veined, ovate to broadly ovate, 15–45 mm, base mostly truncate to shallowly cordate, margins sharply dentate-serrate, apex obtuse, surfaces hirtellous to softly hirsute, hairs ascending, straight, dull gray, sharp-pointed, thick-walled, eglandular. |
Flowers | herkogamous, 1–8, from medial to distal nodes. |
herkogamous, 1–3(–6), commonly solitary or from distal nodes. |
Styles | glabrous. |
sparsely hirtellous. |
Corollas | yellow, throat with fine red to blackish or brown lines extending onto lobes, red to brown dots in throat and lobes present or absent, bilaterally or nearly radially symmetric, bilabiate or nearly regular; tube-throat narrowly funnelform, 11–18 mm, exserted beyond calyx margin; lobes oblong-obovate, apex rounded to notched. |
yellow, red-spotted, bilaterally symmetric, bilabiate; tube-throat narrowly funnelform to broadly cylindric, 13–20 mm, exserted beyond calyx margin; limb expanded 12–22 mm. |
Fruiting pedicels | (7–)10–25 mm, glabrate to glandular-villous. |
(10–)25–75 mm, glabrous or puberulent proximally, hairs stipitate-glandular. |
Fruiting calyces | ridge- to wing-angled, campanulate to cylindric-campanulate, weakly or not inflated, 6–13 mm, villous to glandular-villous, lobes erect to spreading-recurving, strongly unequal to subequal, triangular to linear-lanceolate or narrowly triangular-acuminate, 2–4 mm, apex acute to obtuse. |
sometimes purple-spotted, broadly cylindric-campanulate, inflated, sagittally compressed, 11–15 mm, glabrous, throat not closing, proximal lobe pair slightly upcurving. |
Capsules | included, 6–8 mm. |
included, stipitate, 7–10 mm. |
Anthers | included, glabrous or slightly hirtellous to scabrous. |
included, glabrous. |
2n | = 32. |
= 48, 56. |
Erythranthe moschata |
Erythranthe corallina |
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Phenology | Flowering May–Aug. | Flowering (May–)Jun–Aug. |
Habitat | Springs and seeps, creek edges, moist meadows, ditches, along trails, roadsides, rocky ridges, granite outcrops, shaded and wet places in sagebrush, aspen, fir, spruce-fir, lodgepole pine forests, meadows. | Creek banks, moraine water courses, bogs, marshes, wet meadows, roadside ditches. |
Elevation | (300–)400–3100 m. ((1000–)1300–10200 ft.) | (1400–)1700–2700(–3000) m. ((4600–)5600–8900(–9800) ft.) |
Distribution |
CA; CO; CT; ID; MA; ME; MI; MT; NH; NJ; NV; NY; OR; PA; RI; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM [Introduced in South America (Chile), Europe, e Asia (Japan), Pacific Islands (New Zealand), Australia]
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CA; NV |
Discussion | Earlier segregation of Erythranthe moniliformis as distinct from E. moschata (for example, G. L. Nesom 2012g) emphasized a primarily erect habit and tendency toward sessile to subsessile and more densely arranged cauline leaves in E. moniliformis versus a decumbent to procumbent habit and consistently petiolate leaves on longer internodes in E. moschata. Discontinuities in morphology, geography, and ecology were not confirmed in later study by Nesom (2017). Rhizomes with small, tuberlike swellings can be observed over the whole moschata/moniliformis range, and there apparently are no consistent distinctions in vestiture and corolla size. Mimulus acutidens Reiche (1911), a later homonym of M. acutidens Greene, pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe corallina is a morphologically consistent entity that occurs only in the Sierra Nevada of California and adjacent Nevada (Washoe County and Carson City). Its chromosome number is reported as 2n = 48 and 56, compared to 2n = 28 and 56 in E. tilingii; identities of the E. corallina vouchers should be rechecked and additional counts made, since the occurrence of such wide dysploidy seems unlikely. Compared to the leaf blades of E. tilingii in the strict sense, those of E. corallina are relatively broader (broadly ovate to orbicular-ovate), the plants generally taller, and long-pedicellate flowers occasionally are produced from mid stem or even proximal nodes. The hirsutulous to hirsute vestiture of eglandular hairs on both leaf surfaces is a reliably diagnostic feature and usually easily observed with a 10× lens. Some plants of Erythranthe corallina from San Bernardino County, California, produce decumbent-ascending stems (4–10 cm) and ovate-triangular leaves (blade 5–10 × 3–6 mm), but the dense system of filiform rhizomes, flowers one to three, and hirtellous foliar vestiture serve to identify them. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 401. | FNA vol. 17, p. 410. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus, E. inodora, E. moniliformis, M. crinitus, M. guttatus var. moschatus, M. inodorus, M. leibergii, M. macranthus, M. moniliformis, M. moschatus var. longiflorus, M. moschatus var. moniliformis, M. moschatus var. pallidiflorus | Mimulus corallinus, M. minusculus, M. tilingii var. corallinus |
Name authority | (Douglas ex Lindley) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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