Erythranthe moschata |
Erythranthe cinnabarina |
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mimule musqué, musk monkeyflower, musk-flower, musk-plant, sticky monkey-flower |
Arizona big red monkeyflower |
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Habit | Perennials, rhizomatous, rooting at proximal nodes. | Perennials, rhizomatous. |
Stems | erect, sometimes ascending to decumbent, simple or branched, (2–)5–20 cm, nodes 2–4(or 5), glabrate to glandular-villous, hairs 0.5–2 mm, gland-tipped, internodes evident. |
usually erect to ascending, freely branched, 25–60 cm, glabrous. |
Leaves | usually cauline, basal not persistent, distinctly separated; petiole 0 mm or (0.5–)1–5(–10) mm; blade pinnately veined, oblong-ovate to ovate, (10–)15–40(–50) × 5–25 mm, base obtuse-cuneate to truncate, rounded or subcordate, subclasping to sessile, margins coarsely serrate-dentate to denticulate or subentire, apex acute to obtuse, surfaces glabrate to glandular-villous. |
usually cauline; petiole 0 mm; blade palmately veined, elliptic to oblong-elliptic, elliptic-lanceolate, or broadly lanceolate, 60–125 × 25–46 mm, base narrowly auriculate, clasping to subclasping, margins shallowly dentate, teeth sharp-pointed, apex acute, adaxial surface glabrous or minutely sessile- or stipitate-glandular along veins, lamina glabrous. |
Flowers | herkogamous, 1–8, from medial to distal nodes. |
herkogamous, 2–4(–8), axillary at leafy distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine red to blackish or brown lines extending onto lobes, red to brown dots in throat and lobes present or absent, bilaterally or nearly radially symmetric, bilabiate or nearly regular; tube-throat narrowly funnelform, 11–18 mm, exserted beyond calyx margin; lobes oblong-obovate, apex rounded to notched. |
deep orange, dull orange, red-orange, or deep scarlet, throat yellow-orange, dark red stripes leading onto basal part of lobes, not spotted, palate ridges red, bilaterally symmetric, strongly bilabiate; tube-throat tubular, 29–36 mm, exserted 7–12 mm beyond calyx margin; throat open, palate ridges densely short-villous, hairs yellowish. |
Fruiting pedicels | (7–)10–25 mm, glabrate to glandular-villous. |
50–95 mm. |
Fruiting calyces | ridge- to wing-angled, campanulate to cylindric-campanulate, weakly or not inflated, 6–13 mm, villous to glandular-villous, lobes erect to spreading-recurving, strongly unequal to subequal, triangular to linear-lanceolate or narrowly triangular-acuminate, 2–4 mm, apex acute to obtuse. |
cylindric-campanulate, not inflated, (27–)29–34 mm, minutely stipitate- or sessile-glandular, lobes 7–10 mm, ovate, apex abruptly attenuate to linear-caudate. |
Capsules | included, 6–8 mm. |
included, 14–18 mm. |
Anthers | included, glabrous or slightly hirtellous to scabrous. |
exserted, white-villous, thecae spreading. |
2n | = 32. |
= 16. |
Erythranthe moschata |
Erythranthe cinnabarina |
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Phenology | Flowering May–Aug. | Flowering Jun–Aug(–Sep). |
Habitat | Springs and seeps, creek edges, moist meadows, ditches, along trails, roadsides, rocky ridges, granite outcrops, shaded and wet places in sagebrush, aspen, fir, spruce-fir, lodgepole pine forests, meadows. | Canyons, ravines, streambeds and margins, riparian vegetation, mixed conifer forest. |
Elevation | (300–)400–3100 m. ((1000–)1300–10200 ft.) | 2100–3300 m. (6900–10800 ft.) |
Distribution |
CA; CO; CT; ID; MA; ME; MI; MT; NH; NJ; NV; NY; OR; PA; RI; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM [Introduced in South America (Chile), Europe, e Asia (Japan), Pacific Islands (New Zealand), Australia]
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AZ |
Discussion | Earlier segregation of Erythranthe moniliformis as distinct from E. moschata (for example, G. L. Nesom 2012g) emphasized a primarily erect habit and tendency toward sessile to subsessile and more densely arranged cauline leaves in E. moniliformis versus a decumbent to procumbent habit and consistently petiolate leaves on longer internodes in E. moschata. Discontinuities in morphology, geography, and ecology were not confirmed in later study by Nesom (2017). Rhizomes with small, tuberlike swellings can be observed over the whole moschata/moniliformis range, and there apparently are no consistent distinctions in vestiture and corolla size. Mimulus acutidens Reiche (1911), a later homonym of M. acutidens Greene, pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe cinnabarina is similar to typical E. cardinalis in its spreading anther thecae, relatively short-exserted corolla tube, and its reflexing corolla lobes but distinct in its generally larger leaves with reduced vestiture, fewer flowers, larger calyx and corolla, apically caudate calyx lobes, and its separate geographical range. Erythranthe cinnabarina occurs in Cochise County (Chiricahua Mountains), Graham County (Pinaleño Mountains), and Pima County (Santa Catalina Mountains). Erythranthe verbenacea, with which it sometimes has been confused, occurs at lower elevations (350–2600 m) and ranges over most of the state (Apache, Cochise, Coconino, Gila, Graham, La Paz, Maricopa, Mohave, Pima, Pinal, Santa Cruz, and Yavapai counties). Erythranthe cinnabarina apparently occurs alone (without E. verbenacea) in the Pinaleño Mountains and in the Chiricahua Mountains, but both species have been abundantly documented in the Santa Catalina Mountains, where they sometimes closely co-occur in areas of elevational overlap (for example, at Marshall Gulch, about 2500 m; at Bear Wallow Campground, about 2600 m). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 401. | FNA vol. 17, p. 393. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus, E. inodora, E. moniliformis, M. crinitus, M. guttatus var. moschatus, M. inodorus, M. leibergii, M. macranthus, M. moniliformis, M. moschatus var. longiflorus, M. moschatus var. moniliformis, M. moschatus var. pallidiflorus | |
Name authority | (Douglas ex Lindley) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | G. L. Nesom: Phytoneuron 2014-31: 16, figs. 16, 17. (2014) |
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