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mimule musqué, musk monkeyflower, musk-flower, musk-plant, sticky monkey-flower

Charleston Mountains monkeyflower

Habit Perennials, rhizomatous, rooting at proximal nodes. Annuals, fibrous-rooted, usually dark purplish.
Stems

erect, sometimes ascending to decumbent, simple or branched, (2–)5–20 cm, nodes 2–4(or 5), glabrate to glandular-villous, hairs 0.5–2 mm, gland-tipped, internodes evident.

erect, simple, weakly 4-angled, 4–16(–24) cm, slender, glabrous or sparsely villous-glandular near nodes.

Leaves

usually cauline, basal not persistent, distinctly separated;

petiole 0 mm or (0.5–)1–5(–10) mm;

blade pinnately veined, oblong-ovate to ovate, (10–)15–40(–50) × 5–25 mm, base obtuse-cuneate to truncate, rounded or subcordate, subclasping to sessile, margins coarsely serrate-dentate to denticulate or subentire, apex acute to obtuse, surfaces glabrate to glandular-villous.

basal and cauline, basal usually persistent, largest at mid stem or above, cauline relatively few on long internodes;

petiole: proximals to distals 1–4 mm;

blade palmately 3-veined, ovate to ovate-lanceolate, 5–16(–20) × 3–11 mm, base truncate to subcordate, margins shallowly, evenly crenulate to serrate-dentate or denticulate, apex acute to obtuse, surfaces: proximals glabrous or sparsely villous, distals and bracteals sparsely hirtellous or glabrous abaxially or along distal margin, sparsely villous adaxially, hairs vitreous, flattened, eglandular.

Flowers

herkogamous, 1–8, from medial to distal nodes.

plesiogamous, (1–)4–7, usually from all nodes, usually beginning about mid stem, cleistogamous.

Styles

glabrous.

glabrous.

Corollas

yellow, throat with fine red to blackish or brown lines extending onto lobes, red to brown dots in throat and lobes present or absent, bilaterally or nearly radially symmetric, bilabiate or nearly regular;

tube-throat narrowly funnelform, 11–18 mm, exserted beyond calyx margin;

lobes oblong-obovate, apex rounded to notched.

yellow, sparsely red-dotted, bilaterally or nearly radially symmetric, bilabiate or nearly regular;

tube-throat narrowly cylindric, 4–6 mm, exserted 0.5–1 mm beyond calyx margin;

limb barely expanded.

Fruiting pedicels

(7–)10–25 mm, glabrate to glandular-villous.

deflexed 45–90º at calyx, 6–19 mm, longer than subtending leaves, glabrous or sparsely villous-glandular.

Fruiting calyces

ridge- to wing-angled, campanulate to cylindric-campanulate, weakly or not inflated, 6–13 mm, villous to glandular-villous, lobes erect to spreading-recurving, strongly unequal to subequal, triangular to linear-lanceolate or narrowly triangular-acuminate, 2–4 mm, apex acute to obtuse.

sometimes purple-dotted, broadly elliptic-ovoid, inflated, sagittally compressed, 10–13 mm, minutely hirtellous, sometimes also sparsely glandular, throat closing, adaxial lobe not distinctly longer than abaxial, not falcate.

Capsules

included, 6–8 mm.

included, stipitate, 6–8 mm.

Anthers

included, glabrous or slightly hirtellous to scabrous.

included, glabrous.

2n

= 32.

Erythranthe moschata

Erythranthe charlestonensis

Phenology Flowering May–Aug. Flowering Apr–Jul(–Aug).
Habitat Springs and seeps, creek edges, moist meadows, ditches, along trails, roadsides, rocky ridges, granite outcrops, shaded and wet places in sagebrush, aspen, fir, spruce-fir, lodgepole pine forests, meadows. Grassy slopes, damp soils, moist rocks.
Elevation (300–)400–3100 m. ((1000–)1300–10200 ft.) (900–)1700–2400(–2800) m. ((3000–)5600–7900(–9200) ft.)
Distribution
from FNA
CA; CO; CT; ID; MA; ME; MI; MT; NH; NJ; NV; NY; OR; PA; RI; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM [Introduced in South America (Chile), Europe, e Asia (Japan), Pacific Islands (New Zealand), Australia]
[WildflowerSearch map]
from FNA
AZ; NV
Discussion

Earlier segregation of Erythranthe moniliformis as distinct from E. moschata (for example, G. L. Nesom 2012g) emphasized a primarily erect habit and tendency toward sessile to subsessile and more densely arranged cauline leaves in E. moniliformis versus a decumbent to procumbent habit and consistently petiolate leaves on longer internodes in E. moschata. Discontinuities in morphology, geography, and ecology were not confirmed in later study by Nesom (2017). Rhizomes with small, tuberlike swellings can be observed over the whole moschata/moniliformis range, and there apparently are no consistent distinctions in vestiture and corolla size.

Mimulus acutidens Reiche (1911), a later homonym of M. acutidens Greene, pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe charlestonensis is characterized by its annual duration, autogamous reproduction, small stature, commonly purplish color, regularly ovate, short-petiolate leaves with shallowly crenate margins and (distal leaves) sparsely villous or glabrous adaxial surfaces, hirtellous calyces without a prominently longer adaxial lobe, and lack of glandular hairs. It is endemic to the Charleston (Spring) Mountains of Clark County, Nevada, and one locality (Union Pass) in Mohave County, Arizona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 401. FNA vol. 17, p. 423.
Parent taxa Phrymaceae > Erythranthe Phrymaceae > Erythranthe
Sibling taxa
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms Mimulus moschatus, E. inodora, E. moniliformis, M. crinitus, M. guttatus var. moschatus, M. inodorus, M. leibergii, M. macranthus, M. moniliformis, M. moschatus var. longiflorus, M. moschatus var. moniliformis, M. moschatus var. pallidiflorus
Name authority (Douglas ex Lindley) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) G. L. Nesom: Phytoneuron 2012-40: 80. (2012)
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