Erythranthe moschata |
Erythranthe calciphila |
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mimule musqué, musk monkeyflower, musk-flower, musk-plant, sticky monkey-flower |
Mexican moss monkeyflower |
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Habit | Perennials, rhizomatous, rooting at proximal nodes. | Annuals, usually fibrous-rooted, rarely rooting at proximal nodes. |
Stems | erect, sometimes ascending to decumbent, simple or branched, (2–)5–20 cm, nodes 2–4(or 5), glabrate to glandular-villous, hairs 0.5–2 mm, gland-tipped, internodes evident. |
erect, sometimes decumbent-ascending, branched, 4–30 cm, minutely stipitate-glandular, also delicately villosulous-glandular along whole length. |
Leaves | usually cauline, basal not persistent, distinctly separated; petiole 0 mm or (0.5–)1–5(–10) mm; blade pinnately veined, oblong-ovate to ovate, (10–)15–40(–50) × 5–25 mm, base obtuse-cuneate to truncate, rounded or subcordate, subclasping to sessile, margins coarsely serrate-dentate to denticulate or subentire, apex acute to obtuse, surfaces glabrate to glandular-villous. |
basal and cauline; petiole 1–3 mm, cauline blade slightly or hardly reduced in size from basal, becoming subsessile to sessile (1–3 pairs of cauline leaves); blade palmately 3(–5)-veined, orbicular-ovate to oblong-ovate, 7–28 × 5–22 mm, base truncate to shallowly cuneate, margins shallowly dentate to denticulate, teeth 3–6 per side, apex rounded to obtuse, surfaces villous, hairs thin-walled, vitreous, eglandular or minutely gland-tipped, usually also minutely stipitate-glandular. |
Flowers | herkogamous, 1–8, from medial to distal nodes. |
plesiogamous, 1–6(–10), axillary at all nodes, chasmogamous. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat with fine red to blackish or brown lines extending onto lobes, red to brown dots in throat and lobes present or absent, bilaterally or nearly radially symmetric, bilabiate or nearly regular; tube-throat narrowly funnelform, 11–18 mm, exserted beyond calyx margin; lobes oblong-obovate, apex rounded to notched. |
light yellow, red-spotted, weakly bilaterally symmetric, ± bilabiate; tube-throat narrowly funnelform, 5–7 mm, exserted 1–2 mm beyond calyx margin; limb expanded 2–4 mm. |
Fruiting pedicels | (7–)10–25 mm, glabrate to glandular-villous. |
15–30(–55) mm, minutely stipitate-glandular, sometimes minutely hirtellous and minutely stipitate-glandular, sometimes short glandular-villous. |
Fruiting calyces | ridge- to wing-angled, campanulate to cylindric-campanulate, weakly or not inflated, 6–13 mm, villous to glandular-villous, lobes erect to spreading-recurving, strongly unequal to subequal, triangular to linear-lanceolate or narrowly triangular-acuminate, 2–4 mm, apex acute to obtuse. |
nodding 90º at maturity, usually 3(–5)-veined, ovoid, inflated, sagittally compressed, 6–10 mm, minutely stipitate-glandular, throat closing, lobes usually 3 or 3 and 5 on same plant, if 5 then with 2 interpolated lobes much smaller than abaxial pair. |
Capsules | included, 6–8 mm. |
included, 2–4 mm. |
Anthers | included, glabrous or slightly hirtellous to scabrous. |
included, glabrous. |
2n | = 32. |
= 30, 32. |
Erythranthe moschata |
Erythranthe calciphila |
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Phenology | Flowering May–Aug. | Flowering Mar–Sep(–Nov). |
Habitat | Springs and seeps, creek edges, moist meadows, ditches, along trails, roadsides, rocky ridges, granite outcrops, shaded and wet places in sagebrush, aspen, fir, spruce-fir, lodgepole pine forests, meadows. | Rocky knobs, moist boulders, wet rock faces, roadcuts, seepages, springs, with moss, usually in pine or pine-oak woods. |
Elevation | (300–)400–3100 m. ((1000–)1300–10200 ft.) | 1800–2500 m. (5900–8200 ft.) |
Distribution |
CA; CO; CT; ID; MA; ME; MI; MT; NH; NJ; NV; NY; OR; PA; RI; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM [Introduced in South America (Chile), Europe, e Asia (Japan), Pacific Islands (New Zealand), Australia]
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AZ; Mexico (Chihuahua, Durango, Sinaloa, Sonora) |
Discussion | Earlier segregation of Erythranthe moniliformis as distinct from E. moschata (for example, G. L. Nesom 2012g) emphasized a primarily erect habit and tendency toward sessile to subsessile and more densely arranged cauline leaves in E. moniliformis versus a decumbent to procumbent habit and consistently petiolate leaves on longer internodes in E. moschata. Discontinuities in morphology, geography, and ecology were not confirmed in later study by Nesom (2017). Rhizomes with small, tuberlike swellings can be observed over the whole moschata/moniliformis range, and there apparently are no consistent distinctions in vestiture and corolla size. Mimulus acutidens Reiche (1911), a later homonym of M. acutidens Greene, pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe calciphila is recognized by its annual duration (fibrous-rooted), short, erect stems with few, even-sized leaves (the basal often persistent), delicate stipitate-glandular vestiture, three-lobed calyces that are relatively large in fruit, small corollas, and autogamous reproduction. Plants rarely root at proximal nodes. In Arizona, at the northernmost extremity of its range, E. calciphila occurs in the Chiricahua, Huachuca, and Mule mountains of Cochise County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 401. | FNA vol. 17, p. 424. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus moschatus, E. inodora, E. moniliformis, M. crinitus, M. guttatus var. moschatus, M. inodorus, M. leibergii, M. macranthus, M. moniliformis, M. moschatus var. longiflorus, M. moschatus var. moniliformis, M. moschatus var. pallidiflorus | Mimulus calciphilus, M. minutiflorus |
Name authority | (Douglas ex Lindley) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | (Gentry) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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