Erythranthe linearifolia |
Erythranthe michiganensis |
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primrose monkeyflower, threadleaf primrose monkeyflower |
Michigan monkeyflower |
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Habit | Perennials, rhizomatous, densely cespitose, forming large patches and turfs 0.3–1 m diam. | Perennials, rhizomatous, usually producing numerous leafy stolons from basal nodes, rooting at distal nodes, sometimes forming mats. |
Stems | erect to ascending, simple, 2–10 cm, sparsely hirsute and stipitate-glandular, internodes shortened. |
ascending-erect or basally decumbent, becoming erect in inflorescence, branched, 12–50(–70) cm, glabrous or minutely hirtellous and stipitate-glandular. |
Leaves | basal or near basal, sometimes proximal cauline, subrosulate; petiole 0 mm; blade 1-veined or palmately 3-veined, linear to narrowly oblanceolate, 15–50 × 1.5–5 mm, base long-cuneate, often subclasping, margins entire, dentate-serrate, or distally dentate, apex acute, surfaces glabrous or adaxial sparsely short-pilose, eglandular. |
cauline, basal not persistent; petiole 1–5(–15) mm, mid stem and distals 0 mm; blade palmately 3–5-veined, broadly ovate to broadly ovate-elliptic or suborbicular, 8–30 mm, relatively even-sized or diminishing in size distally, bracteal reduced, slightly falcate, base truncate to cuneate, margins evenly or unevenly dentate-serrate to dentate, teeth 3–8 per side, apex usually rounded, surfaces glabrous or minutely hirtellous and stipitate-glandular. |
Flowers | herkogamous, 1. |
herkogamous, 2–14, mostly from distal nodes or medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, red-spotted or -striped, bilaterally symmetric, weakly bilabiate, loosely hirsute on abaxial side of opening; tube-throat narrowly campanulate, 18–22 mm, exserted beyond calyx margin; lobes broadly obovate-oblong, apex rounded- or truncate-notched, throat open. |
yellow, sometimes faintly red-spotted, bilaterally symmetric, strongly bilabiate; tube-throat cylindric-campanulate, 10–14 mm, exserted 5–8 mm beyond calyx margin; limb expanded 10–15 mm. |
Fruiting pedicels | (40–)65–85(–120) mm, glabrous or sparsely stipitate-glandular near base. |
10–25 mm, villous-glandular to minutely villosulous-glandular. |
Fruiting calyces | winged- or plicate-angled, tubular-campanulate, weakly or not inflated, 9–10(–12) mm, glabrous. |
cylindric-campanulate, inflated, sagittally compressed, 7–10 mm, puberulous to softly hirtellous, mixed with longer, stipitate-glandular hairs, throat not closing, adaxial lobe 2 times longer than others, slightly upcurving. |
Capsules | included, 6–7 mm. |
unknown. |
Anthers | included or slightly exserted, margins ciliate, glabrous. |
included, glabrous. |
2n | = (28)30. |
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Erythranthe linearifolia |
Erythranthe michiganensis |
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Phenology | Flowering Jul–Sep. | Flowering Jun–Aug(–Oct). |
Habitat | Wet banks, Darlingtonia seeps and bogs, seepages in serpentine talus. | Cold calcareous springs, seeps, depressions, streams, alkaline shorelines at mouths of small drainages, steep moraine slopes, bluff bases, commonly in northern white cedar swamps. |
Elevation | 600–2800 m. (2000–9200 ft.) | 500–600 m. (1600–2000 ft.) |
Distribution |
CA
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MI |
Discussion | Erythranthe linearifolia is endemic to serpentine substrates in Shasta, Siskiyou, and Trinity counties; typical E. primuloides occurs in the same area but not on serpentine. Erythranthe linearifolia is distinct from E. primuloides especially in its narrow leaves and cespitose habit. A collection from Tulare County appears to be E. linearifolia (Shevock 10597, CAS), but this appears to be far out of range and the voucher should be reexamined; it probably is better identified as an unusual collection of E. primuloides. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Based on data from allozyme and RAPD studies, morphology, and crossing studies, A. L. Posto and L. A. Prather (2003) raised Erythranthe [Mimulus] michiganensis to specific rank. The species is endemic to a small area in the Mackinac Straits and Grand Traverse regions of Michigan; it is known to be extant at 15 sites and apparently is extinct at three previously known sites. Plants of all but one of the populations are essentially pollen-sterile and reproduce through rhizomes. However, the didynamous stamens and stigma positioned above the adaxial anther pair, along with the relatively large corollas with broadly expanded limb, are reflective of an allogamous breeding system, which seems unusual in view of its apparent sterility. The narrow geographic distribution of E. michiganensis lies within the wider range of E. geyeri, and the two are known to co-occur at two sites, apparently without hybridization or morphologically intermediate individuals. Erythranthe michiganensis (as Mimulus michiganensis) is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 390. | FNA vol. 17, p. 407. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus primuloides var. linearifolius, M. linearifolius, M. primuloides subsp. linearifolius | Mimulus glabratus subsp. michiganensis, M. glabratus var. michiganensis, M. michiganensis |
Name authority | (A. L. Grant) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 35. (2012) | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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