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great purple monkey-flower, great purple or Lewis' monkeyflower, Lewis' monkey flower, pink monkey-flower, purple monkey-flower

John Day or Washington monkeyflower, Washington monkey-flower

Habit Perennials, rhizomatous. Annuals, fibrous-rooted or filiform-taprooted.
Stems

erect, usually simple, (15–)25–60(–75) cm, stipitate-glandular to glandular-villous.

erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

Leaves

cauline;

petiole 0 mm;

blade palmately veined, elliptic to ovate, ovate-lanceolate, or broadly oblanceolate, (10–)25–75(–90) × 5–35 mm, base rounded to cuneate, subclasping, margins denticulate, subentire, or entire, apex acute, surfaces stipitate-glandular to glandular-villous.

cauline, basal not persistent;

petiole 2–14 mm;

blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

Flowers

herkogamous, 2–6(–10), axillary at leafy medial to distal nodes.

herkogamous, 1–6, from proximal to distal nodes.

Styles

glabrous.

hispid-hirtellous.

Corollas

purple, rarely crimson, pale violet, white, pinkish white, yellowish white, or lavender, sometimes lined with red dots, bilaterally symmetric, strongly bilabiate;

tube-throat funnelform, 22–28 mm, exserted beyond calyx margins;

lobe apex usually truncate to shallowly convex, shallowly retuse, throat open.

yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate;

tube-throat funnelform, 8–10 mm, exserted beyond calyx margin;

limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.

Fruiting pedicels

(25–)35–70 mm, stipitate-glandular to glandular-villous.

divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.

Fruiting calyces

broadly cylindric-campanulate, not inflated, 15–22 mm, stipitate-glandular to glandular-villous, tube 12–15(–17) × 9–12 mm.

greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.

Capsules

included, 6–11 mm.

included, 5–8.5 mm.

Anthers

included, white-villous, thecae spreading.

included, glabrous.

2n

= 16.

= 32.

Erythranthe lewisii

Erythranthe washingtonensis

Phenology Flowering Jun–Sep. Flowering May–Sep.
Habitat Stream banks, springs, wet meadows, subalpine slopes, talus, crevices, ditches. Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.
Elevation 600–2900(–3200) m. (2000–9500(–10500) ft.) 700–1300 m. (2300–4300 ft.)
Distribution
from FNA
AK; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
from FNA
OR
[WildflowerSearch map]
Discussion

Erythranthe lewisii in California occurs in Modoc, Siskiyou, and Trinity counties. California records from Shasta County and south previously identified as E. lewisii are identified here as E. erubescens. The record for E. lewisii in Alaska is documented by this collection: Hyder [noted on handwritten label as "New to Alaska, Standley"], damp land, 27 June 1924, K. Whited 1291 (MO).

Apparent exceptions to the characteristic flower color are these: white to lavender in Nevada (Clark County, Charleston Mountains, Train 2068, MO); pinkish white in Washington (Skamania and Yakima counties, Mt. Paddo, Suksdorf 5779, MO); white or tinged with yellow, in Wyoming (Teton County, as described by Nelson in the protologue of Mimulus lewisii var. tetonensis).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 392. FNA vol. 17, p. 396.
Parent taxa Phrymaceae > Erythranthe Phrymaceae > Erythranthe
Sibling taxa
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii
Synonyms Mimulus lewisii, M. lewisii var. tetonensis Mimulus washingtonensis
Name authority (Pursh) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012) (Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)
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