Erythranthe lewisii |
Erythranthe grandis |
|
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great purple monkey-flower, great purple or Lewis' monkeyflower, Lewis' monkey flower, pink monkey-flower, purple monkey-flower |
large monkey-flower, magnificent monkeyflower, magnificent seep monkeyflower |
|
Habit | Perennials, rhizomatous. | Perennials, rhizomatous, sometimes rooting at proximal nodes. |
Stems | erect, usually simple, (15–)25–60(–75) cm, stipitate-glandular to glandular-villous. |
erect, sometimes decumbent basally, branched, often fistulose, (25–)50–120(–160) cm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs. |
Leaves | cauline; petiole 0 mm; blade palmately veined, elliptic to ovate, ovate-lanceolate, or broadly oblanceolate, (10–)25–75(–90) × 5–35 mm, base rounded to cuneate, subclasping, margins denticulate, subentire, or entire, apex acute, surfaces stipitate-glandular to glandular-villous. |
basal and cauline, basal usually not persistent, bracteate in inflorescence; petiole 10–80 mm, gradually reduced distally; blade subpinnately, sometimes palmately, 5–7-veined, ovate to broadly elliptic, 25–60 × 20–40(–60) mm, usually 1–2 times longer than wide, base truncate or truncate-cuneate to subcordate, margins crenulate to dentate, proximally sometimes sublyrate, apex rounded to obtuse, surfaces of distals densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs. |
Flowers | herkogamous, 2–6(–10), axillary at leafy medial to distal nodes. |
herkogamous, 8–26, mostly from distal nodes, usually in bracteate racemes. |
Styles | glabrous. |
hirtellous. |
Corollas | purple, rarely crimson, pale violet, white, pinkish white, yellowish white, or lavender, sometimes lined with red dots, bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 22–28 mm, exserted beyond calyx margins; lobe apex usually truncate to shallowly convex, shallowly retuse, throat open. |
yellow, red-dotted within, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, (14–)16–24 mm, exserted (8–)10–15 mm beyond calyx margin; limb broadly expanded. |
Fruiting pedicels | (25–)35–70 mm, stipitate-glandular to glandular-villous. |
10–35 mm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs. |
Fruiting calyces | broadly cylindric-campanulate, not inflated, 15–22 mm, stipitate-glandular to glandular-villous, tube 12–15(–17) × 9–12 mm. |
straight-erect or nodding 45–100º, ovate-campanulate, inflated, sagittally compressed, 15–22(–25) mm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs, throat closing. |
Capsules | included, 6–11 mm. |
included, 8–12 mm. |
Anthers | included, white-villous, thecae spreading. |
included, glabrous. |
2n | = 16. |
= 28. |
Erythranthe lewisii |
Erythranthe grandis |
|
Phenology | Flowering Jun–Sep. | Flowering (Apr–)May–Jul(–Sep). |
Habitat | Stream banks, springs, wet meadows, subalpine slopes, talus, crevices, ditches. | Beaches, dunes, coastal bluffs, wet cliff faces, mud flats and seeps, marshes, drainage ditches, creeks, rarely in coastal sage scrub. |
Elevation | 600–2900(–3200) m. (2000–9500(–10500) ft.) | 0–200(–800) m. (0–700(–2600) ft.) |
Distribution |
AK; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
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CA; OR
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Discussion | Erythranthe lewisii in California occurs in Modoc, Siskiyou, and Trinity counties. California records from Shasta County and south previously identified as E. lewisii are identified here as E. erubescens. The record for E. lewisii in Alaska is documented by this collection: Hyder [noted on handwritten label as "New to Alaska, Standley"], damp land, 27 June 1924, K. Whited 1291 (MO). Apparent exceptions to the characteristic flower color are these: white to lavender in Nevada (Clark County, Charleston Mountains, Train 2068, MO); pinkish white in Washington (Skamania and Yakima counties, Mt. Paddo, Suksdorf 5779, MO); white or tinged with yellow, in Wyoming (Teton County, as described by Nelson in the protologue of Mimulus lewisii var. tetonensis). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The densely, evenly puberulent vestiture of pedicels, calyces, and distal stems usually is diagnostic, especially in combination with the large flowers (corollas and mature calyces) and tall stature. Plants from scattered collections are much shorter than normal but have large corollas and characteristic vestiture. Erythranthe grandis characteristically occurs in coastal localities from southern California to northern Oregon but also is found in inland localities and habitats near the coast but well away from salt spray. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 392. | FNA vol. 17, p. 412. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus lewisii, M. lewisii var. tetonensis | Mimulus guttatus var. grandis, M. grandis, M. guttatus subsp. litoralis, M. langsdorffii var. grandis, M. procerus |
Name authority | (Pursh) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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