Erythranthe latidens |
Erythranthe scouleri |
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broad-tooth monkeyflower |
Columbia River monkey-flower, Scouler's monkey-flower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Perennials, rhizomatous, producing long, sparsely leafy runners from basal nodes. |
Stems | ascending to ascending-erect, geniculate at nodes, usually many-branched from base, 3–10(–25) cm, short stipitate-glandular or sessile-glandular, hairs 0.1–0.3 mm, gland-tipped. |
erect, simple or few-branched, 15–80 cm, glabrous. |
Leaves | basal and cauline, largest at base or near mid stem, sometimes unreduced in size up to distalmost nodes; petiole 0 mm; cauline blade palmately 3(–5)-veined, ovate to ovate-lanceolate, 8–26(–35) mm, base abruptly cuneate to rounded, sometimes subauriculate, margins entire or barely mucronulate to shallowly dentate-mucronulate, teeth or mucronulae 1–3 per side, apex acute to rounded, surfaces short stipitate-glandular or sessile-glandular, hairs 0.1–0.3 mm, gland-tipped. |
cauline, basal usually not persistent; petiole: proximals and medial 10–25 mm (gradually merging into blade), distals 0 mm; blade palmately (3–)5–7-veined to subpinnate, (proximal to medial) oblong-elliptic to oblong-lanceolate, 25–60 × 8–18 mm, usually 3–4 times longer than wide, base attenuate, margins evenly, shallowly dentate or crenate to mucronate or mucronulate, teeth 10–20 per side, sometimes more deeply toothed at base, apex obtuse to acute, surfaces glabrous. |
Flowers | plesiogamous, (1–)3–12, from medial to distal nodes. |
herkogamous, (1 or)2–8, from distal nodes. |
Styles | glabrous. |
minutely, prominently hirsutulous to villosulous. |
Corollas | white to pinkish or flesh colored, rarely yellowish, throat and abaxial lobes red-spotted, nearly radially symmetric; tube-throat cylindric, 5–6(–8) mm, exserted beyond calyx margin; limb barely widened, lobes broadly obovate, apex rounded. |
yellow, without red markings, bilaterally symmetric, bilabiate; tube-throat funnelform, 20–24 mm, exserted 10–15 mm beyond calyx margin; limb expanded 22–30 mm. |
Fruiting pedicels | straight, 11–28 mm, short stipitate-glandular or sessile-glandular, hairs 0.1–0.3 mm, gland-tipped. |
20–25 mm, glabrous. |
Fruiting calyces | purplish, prominently 5-angled, tubular-campanulate or ovoid-ellipsoid, strongly inflated, 8–12 mm, margins distinctly toothed or lobed, short stipitate-glandular or sessile-glandular, hairs 0.1–0.3 mm, gland-tipped, lobes pronounced, erect. |
ovoid, inflated, sagittally compressed, 13–14 mm, glabrous, throat closing. |
Capsules | included, 6–7 mm. |
unknown. |
Anthers | included, glabrous. |
included, glabrous. |
Erythranthe latidens |
Erythranthe scouleri |
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Phenology | Flowering Apr–Jun. | Flowering May–Aug. |
Habitat | Drained flats or slopes subject to vernal inundation, depressions in open fields, bare clay soils, vacant lots, roadsides. | Salt marshes, river banks. |
Elevation | 10–800 m. (0–2600 ft.) | 0–20 m. (0–100 ft.) |
Distribution |
CA; OR; Mexico (Baja California)
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OR; WA |
Discussion | The distinction between Erythranthe latidens and E. inflatula sometimes seems arbitrary, perhaps because of gene exchange where they are sympatric in northwestern California. Erythranthe latidens in Oregon is known only from southern Harney County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe scouleri is distinctive in its oblong-elliptic leaves with long-tapering bases and closely toothed margins, completely glabrous vestiture, long, leafy runners from basal cauline nodes, large corollas with broad limbs, and prominently hairy styles. Several features suggest a close relationship to E. decora, particularly its very large corollas, hairy styles, closely toothed leaf margins, tall, simple, and erect stems, numerous runners, and its geographic range. All collections apparently have been made near the mouth of the Columbia River in Clatsop and Columbia counties, Oregon (G. L. Nesom 2013d). Recent observations (Alexander John Wright, pers. comm.; photos http://www.inaturalist.org/observations/3537008) indicate that it also occurs in Wahkiakum County, Washington, in the Julia Butler Hansen Refuge. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 400. | FNA vol. 17, p. 415. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus inconspicuus var. latidens, M. latidens | Mimulus scouleri, M. guttatus subsp. scouleri |
Name authority | (A. Gray) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | (Hooker) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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