Erythranthe latidens |
Erythranthe cordata |
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broad-tooth monkeyflower |
tinytooter monkeyflower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Annuals, fibrous-rooted, sometimes producing leafy runners from basal nodes, stems often rooting at proximal nodes and appearing rhizomelike. |
Stems | ascending to ascending-erect, geniculate at nodes, usually many-branched from base, 3–10(–25) cm, short stipitate-glandular or sessile-glandular, hairs 0.1–0.3 mm, gland-tipped. |
usually erect, usually simple, usually fistulose, 12–40(–100) cm, sparsely stipitate-glandular, hairs fine, gland-tipped. |
Leaves | basal and cauline, largest at base or near mid stem, sometimes unreduced in size up to distalmost nodes; petiole 0 mm; cauline blade palmately 3(–5)-veined, ovate to ovate-lanceolate, 8–26(–35) mm, base abruptly cuneate to rounded, sometimes subauriculate, margins entire or barely mucronulate to shallowly dentate-mucronulate, teeth or mucronulae 1–3 per side, apex acute to rounded, surfaces short stipitate-glandular or sessile-glandular, hairs 0.1–0.3 mm, gland-tipped. |
basal and cauline, basal persistent; petiole: basal and proximals 6–20(–40) mm, midcauline to distals 0 mm; blade not connate, palmately 3–5(–7)-veined, orbicular to broadly elliptic-ovate or oblong-elliptic, cauline becoming broadly ovate to narrowly reniform, basal and mid cauline 15–30(–50) mm, gradually reduced in size distally to 6 mm, basal largest, distal closely paired, auriculate-subclasping, base cuneate to truncate or shallowly cordate, margins shallowly, evenly to unevenly dentate, apex obtuse to rounded, surfaces glabrous. |
Flowers | plesiogamous, (1–)3–12, from medial to distal nodes. |
plesiogamous, (5–)10–16, at distal nodes, in bracteate racemes, chasmogamous or cleistogamous. |
Styles | glabrous. |
glabrous. |
Corollas | white to pinkish or flesh colored, rarely yellowish, throat and abaxial lobes red-spotted, nearly radially symmetric; tube-throat cylindric, 5–6(–8) mm, exserted beyond calyx margin; limb barely widened, lobes broadly obovate, apex rounded. |
yellow, red-spotted, abaxial limb deeper yellow, weakly bilaterally or radially symmetric, weakly bilabiate or regular; tube-throat sometimes tubular and not opening (cleistogamous), 8–14 mm, exserted 1–3 mm beyond calyx margin; limb not expanded or expanded 9–14 mm. |
Fruiting pedicels | straight, 11–28 mm, short stipitate-glandular or sessile-glandular, hairs 0.1–0.3 mm, gland-tipped. |
10–30(–45) mm, longer than subtending leaves, minutely stipitate-glandular. |
Fruiting calyces | purplish, prominently 5-angled, tubular-campanulate or ovoid-ellipsoid, strongly inflated, 8–12 mm, margins distinctly toothed or lobed, short stipitate-glandular or sessile-glandular, hairs 0.1–0.3 mm, gland-tipped, lobes pronounced, erect. |
nodding 45–90º, not red-dotted, broadly elliptic-ovoid, inflated, sagittally compressed, (8–)14–18(–20) mm, glabrous or sparsely stipitate-glandular to hirsutulous, sometimes mixed glandular-hirsutulous, throat closing, adaxial lobe not distinctly longer than abaxial, not falcate. |
Capsules | included, 6–7 mm. |
included, stipitate, 5–7 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 60. |
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Erythranthe latidens |
Erythranthe cordata |
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Phenology | Flowering Apr–Jun. | Flowering (Jan–)Mar–Jun(–Nov). |
Habitat | Drained flats or slopes subject to vernal inundation, depressions in open fields, bare clay soils, vacant lots, roadsides. | Springs, seeps, stream edges, muddy banks, flood plains, marshes and swamps, wash bottoms, wet depressions, wet places among boulders. |
Elevation | 10–800 m. (0–2600 ft.) | (600–)800–2400(–3000) m. ((2000–)2600–7900(–9800) ft.) |
Distribution |
CA; OR; Mexico (Baja California)
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AZ; CA; CO; NM; NV; TX; UT
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Discussion | The distinction between Erythranthe latidens and E. inflatula sometimes seems arbitrary, perhaps because of gene exchange where they are sympatric in northwestern California. Erythranthe latidens in Oregon is known only from southern Harney County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe cordata is characterized by its fibrous-rooted habit (annual in duration, without rhizomes but commonly rooting at the proximal nodes), short corollas and autogamous reproduction (anthers and stigma at the same level), closed calyces, sparsely villous-glandular vestiture (lacking hirtellous, eglandular hairs), and stems commonly fistulose in larger plants. The short corollas and other features of autogamous reproduction of E. cordata are diagnostic and prominent. Plants of E. cordata are highly variable in size, from tiny fibrous-rooted plants with nearly filiform stems to much larger individuals with fistulose stems rooting at proximal nodes. Erythranthe cordata and E. nasuta are sympatric in Arizona, southeastern New Mexico, and southern Utah, and small plants of each species may be similar in aspect, both with cleistogamous flowers and reduced vestiture. Erythranthe nasuta can be recognized by its distal and bracteal leaves with hirtellous to hirsutulous adaxial surfaces; a 10/x lens usually is required to see this feature, and it sometimes is most obvious around the leaf margins. The common name of Erythranthe cordata alludes to a fancied resemblance of the corollas to the horn of a diminutive trumpet. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 400. | FNA vol. 17, p. 422. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus inconspicuus var. latidens, M. latidens | Mimulus cordatus, M. maguirei |
Name authority | (A. Gray) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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