Erythranthe laciniata |
Erythranthe ptilota |
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cut-leaf monkeyflower |
musk monkeyflower, musk-flower, sessile-leaf monkey-flower, wing-leaf monkeyflower |
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Habit | Annuals, slender-taprooted or fibrous-rooted. | Perennials, rhizomatous, sometimes rooting at proximal nodes. |
Stems | erect, simple or branched from base, 3–38 cm, glabrous or sparsely hirtellous, finely villosulous-glandular above nodes. |
prostrate, sometimes decumbent to ascending, few-branched, 20–80 cm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones, internodes evident. |
Leaves | cauline, basal deciduous by flowering; petiole 1–35 mm, distals 0 mm; blade 1-veined or palmately 3-veined, elliptic to elliptic-obovate, oblanceolate, or oblong, 3–55 mm, longer than wide, base attenuate, margins narrowly pinnately lobed or dissected, sometimes merely shallowly toothed, apex acute to obtuse, surfaces glabrate. |
cauline, basal not persistent, often congested; petiole 0 mm, rarely 1–2(–3) mm; blade pinnately veined, oblong-lanceolate, 30–70 × 10–22 mm, base rounded, margins denticulate to dentate, apex acute, surfaces villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
Flowers | plesiogamous, 2–8, from medial to distal nodes, chasmogamous, sometimes cleistogamous. |
herkogamous, 4–10, from medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat red-spotted, abaxial limb of larger usually with 1 large red splotch, bilaterally symmetric, ± bilabiate; tube-throat funnelform, 4–6 mm, exserted 1–2 mm beyond calyx margin; limb expanded 5–6 mm. |
yellow, throat with fine blackish or brownish lines on all sides, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly campanulate, 15–18 mm, exserted beyond calyx margin; lobe apex rounded. |
Fruiting pedicels | nodding 30–140º at calyx base, 5–25 mm. |
(15–)22–50 mm, villous, hairs 1–2 mm, eglandular, sometimes mixed with much shorter stipitate-glandular ones. |
Fruiting calyces | red-spotted, cylindric-campanulate, inflated, sagittally compressed, 8–10 mm, glabrate, throat closing, lobes ca. equal size or adaxial slightly longer. |
wing- or plicate-angled, cylindric-campanulate, weakly inflated, 10–12 mm, villous-glandular, hairs gland-tipped, lobes distinctly spreading, strongly unequal, linear-lanceolate to narrowly triangular, 5–9 mm, apex long acuminate-apiculate. |
Capsules | included, stipitate, 5–7 mm. |
included, 6–8 mm. |
Anthers | included, glabrous. |
included, finely hirtellous to hispidulous. |
2n | = 28. |
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Erythranthe laciniata |
Erythranthe ptilota |
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Phenology | Flowering Apr–Jul(–Aug). | Flowering (May–)Jun–Aug. |
Habitat | Cracks, depressions, and seeps in granite outcrops, ledges, talus and scree, rocky streamsides, rocky slopes, roadsides, intermittent drainages. | Creek banks, gravel bars, flood plains, shallow ditches and natural drainages, swales, damp banks, wet sand, moist soils in coniferous woods, marshes, bogs. |
Elevation | 900–2300(–3300) m. (3000–7500(–10800) ft.) | 0–1000(–1900) m. (0–3300(–6200) ft.) |
Distribution |
CA
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CA; OR; WA; BC
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Discussion | Erythranthe laciniata is known from Amador County south to Kern County. As in Erythranthe nasuta, the adaxial calyx lobe in E. laciniata tends to be narrowly lanceolate to triangular (noselike) and perceptibly falcate, curving slightly upward both in flower and in fruit. The adaxial lobe is not so prominently protruding as it often is in E. nasuta. Corolla size is variable in Erythranthe laciniata, but the size of those with an open throat (versus much reduced in size and apparently cleistogamous) is not strongly correlated with size of the individual plant, and all on one plant are about the same size (compare with E. nasuta). Corollas on some plants, however, are all or nearly all greatly reduced and apparently cleistogamous. Fertilization in even the larger corollas apparently is autogamous; the anther pairs are slightly separated or equal in level, and the stigma is in the middle of the anthers or at the level of the adaxial pair. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe ptilota is recognized by its prostrate to decumbent or decumbent-ascending habit, large, consistently sessile leaves, densely villous vestiture, long pedicels, large calyces and corollas, hispid-hirtellous anthers, and particularly by its long, strongly unequal, linear-triangular calyx lobes usually distally falcate. Leaf bases typically are truncate to rounded or subcordate. Rarely the leaves are short-petiolate, but in such cases, the distinctive leaf bases, vestiture, calyx morphology, and pubescent anthers are diagnostic. Erythranthe ptilota is widely sympatric with E. moschata but usually occurs at lower elevations and characteristically in wetter habitats. The epithet ptilota (Greek ptilotos, winged) alludes to a fancied winglike aspect of the pairs of sessile leaves. A population system of Erythranthe ptilota-like plants occurs in southern California, about 480 km disjunct from the main range of the species. These plants have the prostrate habit, large leaves, long pedicels, and large corollas of E. ptilota, but the calyx lobes are variable in length and usually do not show the characteristic attenuate-apiculate apices. The southern California plants are identified here as E. moschata. Erythranthe ptilota is a new name at specific rank for Mimulus moschatus var. sessilifolius [not E. sessilifolia (Maximowicz) G. L. Nesom]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 419. | FNA vol. 17, p. 402. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus laciniatus, M. eisenii | Mimulus moschatus var. sessilifolius |
Name authority | (A. Gray) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | G. L. Nesom: Phytoneuron 2017-17: 4. (2017) |
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