Erythranthe laciniata |
Erythranthe hymenophylla |
|
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cut-leaf monkeyflower |
thin-sepal monkeyflower |
|
Habit | Annuals, slender-taprooted or fibrous-rooted. | Annuals, filiform-taprooted. |
Stems | erect, simple or branched from base, 3–38 cm, glabrous or sparsely hirtellous, finely villosulous-glandular above nodes. |
prostrate to ascending-erect, sharply bent at basal nodes, simple or few-branched, 5–25 cm, glandular-puberulent to glandular-villous, hairs 0.1–0.8 mm, vitreous, flattened, multicellular, gland-tipped. |
Leaves | cauline, basal deciduous by flowering; petiole 1–35 mm, distals 0 mm; blade 1-veined or palmately 3-veined, elliptic to elliptic-obovate, oblanceolate, or oblong, 3–55 mm, longer than wide, base attenuate, margins narrowly pinnately lobed or dissected, sometimes merely shallowly toothed, apex acute to obtuse, surfaces glabrate. |
basal and cauline, largest at mid stem; petiole 6–30 mm; blade pinnately veined, broadly lanceolate to ovate, 10–35 × 10–30 mm, distinctly membranous, base cuneate to shallowly cordate, margins coarsely dentate to shallowly denticulate or entire, apex acute to obtuse, surfaces glandular-puberulent to glandular-villous, hairs 0.1–0.8 mm, vitreous, flattened, multicellular, gland-tipped, glandular. |
Flowers | plesiogamous, 2–8, from medial to distal nodes, chasmogamous, sometimes cleistogamous. |
herkogamous, 1–6, from proximal to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat red-spotted, abaxial limb of larger usually with 1 large red splotch, bilaterally symmetric, ± bilabiate; tube-throat funnelform, 4–6 mm, exserted 1–2 mm beyond calyx margin; limb expanded 5–6 mm. |
light yellow, throat and abaxial lobes red- or purple-spotted, sometimes with small white patches, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 10–14 mm, exserted beyond calyx margin; lobes obovate-oblong, apex rounded to truncate or notched. |
Fruiting pedicels | nodding 30–140º at calyx base, 5–25 mm. |
divergent at right angles from stem, usually closely paired, 10–45 mm, negatively phototropic, causing capsules to be pressed against a cliff face or crevice at time of dehiscence, glandular-puberulent to glandular-villous, hairs 0.1–0.8 mm, vitreous, flattened, multicellular, gland-tipped. |
Fruiting calyces | red-spotted, cylindric-campanulate, inflated, sagittally compressed, 8–10 mm, glabrate, throat closing, lobes ca. equal size or adaxial slightly longer. |
angled, tubular-campanulate, slightly inflated, 5–7 mm, margins distinctly toothed or lobed, sparsely stipitate-glandular, lobes pronounced, erect. |
Capsules | included, stipitate, 5–7 mm. |
included, 3–6 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28. |
= 32. |
Erythranthe laciniata |
Erythranthe hymenophylla |
|
Phenology | Flowering Apr–Jul(–Aug). | Flowering Apr–Aug(–Sep). |
Habitat | Cracks, depressions, and seeps in granite outcrops, ledges, talus and scree, rocky streamsides, rocky slopes, roadsides, intermittent drainages. | Steep, seasonally moist, basalt cliffs with west or southwest exposure, mesic coniferous forests. |
Elevation | 900–2300(–3300) m. (3000–7500(–10800) ft.) | 800–1300 m. (2600–4300 ft.) |
Distribution |
CA
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ID; MT; OR |
Discussion | Erythranthe laciniata is known from Amador County south to Kern County. As in Erythranthe nasuta, the adaxial calyx lobe in E. laciniata tends to be narrowly lanceolate to triangular (noselike) and perceptibly falcate, curving slightly upward both in flower and in fruit. The adaxial lobe is not so prominently protruding as it often is in E. nasuta. Corolla size is variable in Erythranthe laciniata, but the size of those with an open throat (versus much reduced in size and apparently cleistogamous) is not strongly correlated with size of the individual plant, and all on one plant are about the same size (compare with E. nasuta). Corollas on some plants, however, are all or nearly all greatly reduced and apparently cleistogamous. Fertilization in even the larger corollas apparently is autogamous; the anther pairs are slightly separated or equal in level, and the stigma is in the middle of the anthers or at the level of the adaxial pair. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In the protologue, R. J. Meinke observed that plants of Erythranthe hymenophylla have reflexed fruiting pedicels that increase seed dispersal back onto the vertical cliff wall, the characteristic habitat of the species. The hanging habit of E. hymenophylla is reflected in a sharp (90º to 180º) bend in the basal nodes and the long pedicels that are closely paired and divergent in parallel at about right angles from the stem. The species also is characterized by it very short calyx to corolla length, relatively short capsules, and large seeds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 419. | FNA vol. 17, p. 397. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus laciniatus, M. eisenii | Mimulus hymenophyllus |
Name authority | (A. Gray) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | (Meinke) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) |
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