Erythranthe laciniata |
Erythranthe arenicola |
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cut-leaf monkeyflower |
beach monkeyflower |
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Habit | Annuals, slender-taprooted or fibrous-rooted. | Annuals, fibrous-rooted or slender-taprooted, rarely rooting at nodes. |
Stems | erect, simple or branched from base, 3–38 cm, glabrous or sparsely hirtellous, finely villosulous-glandular above nodes. |
erect, rarely prostrate to prostrate-ascending, few-branched, 3–17 cm, moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular. |
Leaves | cauline, basal deciduous by flowering; petiole 1–35 mm, distals 0 mm; blade 1-veined or palmately 3-veined, elliptic to elliptic-obovate, oblanceolate, or oblong, 3–55 mm, longer than wide, base attenuate, margins narrowly pinnately lobed or dissected, sometimes merely shallowly toothed, apex acute to obtuse, surfaces glabrate. |
basal and cauline; petiole: basal 2–8 mm or mid and distals absent; blade palmately 3–5-veined, suborbicular to broadly ovate or depressed-ovate, 5–17 × 6–15 mm, base truncate or truncate-cuneate to subcordate, margins subentire or crenulate, apex rounded to obtuse, surfaces moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular. |
Flowers | plesiogamous, 2–8, from medial to distal nodes, chasmogamous, sometimes cleistogamous. |
herkogamous, 1–6, at distal nodes, chasmogamous. |
Styles | glabrous. |
hirtellous. |
Corollas | yellow, throat red-spotted, abaxial limb of larger usually with 1 large red splotch, bilaterally symmetric, ± bilabiate; tube-throat funnelform, 4–6 mm, exserted 1–2 mm beyond calyx margin; limb expanded 5–6 mm. |
yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 11–20 mm, exserted 4–8 mm beyond calyx margin; limb expanded 10–18 mm. |
Fruiting pedicels | nodding 30–140º at calyx base, 5–25 mm. |
9–17 mm, moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular. |
Fruiting calyces | red-spotted, cylindric-campanulate, inflated, sagittally compressed, 8–10 mm, glabrate, throat closing, lobes ca. equal size or adaxial slightly longer. |
nodding, ovoid-campanulate, inflated, sagittally compressed, 9–16 mm, moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular, throat closing. |
Capsules | included, stipitate, 5–7 mm. |
included, 5–12 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28. |
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Erythranthe laciniata |
Erythranthe arenicola |
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Phenology | Flowering Apr–Jul(–Aug). | Flowering Apr–Aug. |
Habitat | Cracks, depressions, and seeps in granite outcrops, ledges, talus and scree, rocky streamsides, rocky slopes, roadsides, intermittent drainages. | Sandy beaches, especially in moist hollows among dunes, sea cliff bases, chaparral near beaches, mudstone outcrops. |
Elevation | 900–2300(–3300) m. (3000–7500(–10800) ft.) | 0–100 m. (0–300 ft.) |
Distribution |
CA
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CA |
Discussion | Erythranthe laciniata is known from Amador County south to Kern County. As in Erythranthe nasuta, the adaxial calyx lobe in E. laciniata tends to be narrowly lanceolate to triangular (noselike) and perceptibly falcate, curving slightly upward both in flower and in fruit. The adaxial lobe is not so prominently protruding as it often is in E. nasuta. Corolla size is variable in Erythranthe laciniata, but the size of those with an open throat (versus much reduced in size and apparently cleistogamous) is not strongly correlated with size of the individual plant, and all on one plant are about the same size (compare with E. nasuta). Corollas on some plants, however, are all or nearly all greatly reduced and apparently cleistogamous. Fertilization in even the larger corollas apparently is autogamous; the anther pairs are slightly separated or equal in level, and the stigma is in the middle of the anthers or at the level of the adaxial pair. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
F. W. Pennell (1947, 1951) considered Erythranthe arenicola an endemic of Monterey County, but plants from adjacent San Luis Obispo and Santa Cruz counties also belong here. Most of the localities are at seaside, but some are more than a mile inland. Erythranthe arenicola is hypothesized here to be a derivative of E. guttata or E. grandis, retaining the herkogamous breeding system of its putative ancestor but reduced in size and duration. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 419. | FNA vol. 17, p. 412. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus laciniatus, M. eisenii | Mimulus guttatus subsp. arenicola |
Name authority | (A. Gray) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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