Erythranthe laciniata |
Erythranthe arenaria |
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cut-leaf monkeyflower |
sand-loving monkeyflower |
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Habit | Annuals, slender-taprooted or fibrous-rooted. | Annuals, fibrous-rooted or filiform-taprooted. |
Stems | erect, simple or branched from base, 3–38 cm, glabrous or sparsely hirtellous, finely villosulous-glandular above nodes. |
erect to ascending, straight or geniculate at nodes, simple or branched, 5–20 cm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
Leaves | cauline, basal deciduous by flowering; petiole 1–35 mm, distals 0 mm; blade 1-veined or palmately 3-veined, elliptic to elliptic-obovate, oblanceolate, or oblong, 3–55 mm, longer than wide, base attenuate, margins narrowly pinnately lobed or dissected, sometimes merely shallowly toothed, apex acute to obtuse, surfaces glabrate. |
basal and cauline; petiole 0 mm or proximals 1–3(–5) mm; blade 1-veined or palmately 3-veined, elliptic to narrowly elliptic, ovate-elliptic, or ovate-lanceolate, 5–12(–17) × 3–7 mm, base rounded to cuneate-attenuate, margins entire or sparsely dentate to serrate, apex acuminate to acute or obtuse, surfaces villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
Flowers | plesiogamous, 2–8, from medial to distal nodes, chasmogamous, sometimes cleistogamous. |
herkogamous, 1–22, from proximal to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, throat red-spotted, abaxial limb of larger usually with 1 large red splotch, bilaterally symmetric, ± bilabiate; tube-throat funnelform, 4–6 mm, exserted 1–2 mm beyond calyx margin; limb expanded 5–6 mm. |
yellow, abaxial limb red-dotted, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 9–12(–14) mm, exserted beyond calyx margin; lobes broadly obovate, apex rounded. |
Fruiting pedicels | nodding 30–140º at calyx base, 5–25 mm. |
divergent-arcuate, 10–23 mm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
Fruiting calyces | red-spotted, cylindric-campanulate, inflated, sagittally compressed, 8–10 mm, glabrate, throat closing, lobes ca. equal size or adaxial slightly longer. |
usually red-dotted, narrowly campanulate, not or weakly inflated, 5–7(–9) mm, margins distinctly toothed or lobed, villous-glandular, ribs angled, lobes pronounced, erect. |
Capsules | included, stipitate, 5–7 mm. |
included, 4–7 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28. |
= 32. |
Erythranthe laciniata |
Erythranthe arenaria |
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Phenology | Flowering Apr–Jul(–Aug). | Flowering May–Sep. |
Habitat | Cracks, depressions, and seeps in granite outcrops, ledges, talus and scree, rocky streamsides, rocky slopes, roadsides, intermittent drainages. | Sandy flats, bars, gullies, washes, trails, roadcuts, seasonal creek beds and drainages, rocky slopes, seepy loam, ditches, lake edges, meadows, openings in pine-fir and pine-oak woodlands. |
Elevation | 900–2300(–3300) m. (3000–7500(–10800) ft.) | (100–)500–2600(–2800) m. ((300–)1600–8500(–9200) ft.) |
Distribution |
CA
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CA
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Discussion | Erythranthe laciniata is known from Amador County south to Kern County. As in Erythranthe nasuta, the adaxial calyx lobe in E. laciniata tends to be narrowly lanceolate to triangular (noselike) and perceptibly falcate, curving slightly upward both in flower and in fruit. The adaxial lobe is not so prominently protruding as it often is in E. nasuta. Corolla size is variable in Erythranthe laciniata, but the size of those with an open throat (versus much reduced in size and apparently cleistogamous) is not strongly correlated with size of the individual plant, and all on one plant are about the same size (compare with E. nasuta). Corollas on some plants, however, are all or nearly all greatly reduced and apparently cleistogamous. Fertilization in even the larger corollas apparently is autogamous; the anther pairs are slightly separated or equal in level, and the stigma is in the middle of the anthers or at the level of the adaxial pair. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe arenaria is known from a cluster of six counties of the central Sierra Nevada: Fresno, Madera, Mariposa, Merced, Tulare, and Tuolumne. Most plants of Erythranthe arenaria have relatively even-sized cauline leaves, all sessile to proximally subsessile. Plants in the Yosemite area with persistent basal leaves that are short-petiolate, ovate with a cuneate base, and relatively larger than the more distal cauline ones, and possibly related to E. arenaria, have been named M. floribundus var. subulatus. These might be construed as showing the influence of E. geniculata, but that species occurs only at the lower range of elevation of E. arenaria, while plants referable to Mimulus floribundus var. subulatus occur at least to 2300 m and also have the erect habit characteristic of E. arenaria. These variants should be investigated, especially in the Yosemite area where they appear to be relatively common, with the possibility that they indeed represent a distinct entity. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 419. | FNA vol. 17, p. 403. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus laciniatus, M. eisenii | Mimulus arenarius, M. floribundus var. subulatus, M. multiflorus, M. subulatus, M. trisulcatus |
Name authority | (A. Gray) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) | (A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) |
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