Erythranthe jungermannioides |
Erythranthe grandis |
|
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Jungermann's monkeyflower, liverwort monkey-flower |
large monkey-flower, magnificent monkeyflower, magnificent seep monkeyflower |
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Habit | Perennials, stoloniferous, stolons thin, forming overwintering turions. | Perennials, rhizomatous, sometimes rooting at proximal nodes. |
Stems | decumbent to procumbent, simple or branching near base, 5–38(–60) cm, densely glandular-villous, hairs 0.5–1.2(–1.5) mm, gland-tipped, internodes evident. |
erect, sometimes decumbent basally, branched, often fistulose, (25–)50–120(–160) cm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs. |
Leaves | cauline, basal not persistent; petiole 2–5(–20) mm; blade subpalmately to pinnately veined, broadly ovate to broadly lanceolate, 7–35(–40) × 8–25 mm, base rounded, margins sharply, irregularly dentate to denticulate, apex acute to obtuse, surfaces glandular-villous. |
basal and cauline, basal usually not persistent, bracteate in inflorescence; petiole 10–80 mm, gradually reduced distally; blade subpinnately, sometimes palmately, 5–7-veined, ovate to broadly elliptic, 25–60 × 20–40(–60) mm, usually 1–2 times longer than wide, base truncate or truncate-cuneate to subcordate, margins crenulate to dentate, proximally sometimes sublyrate, apex rounded to obtuse, surfaces of distals densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs. |
Flowers | herkogamous, 2 or 3, from medial to distal nodes. |
herkogamous, 8–26, mostly from distal nodes, usually in bracteate racemes. |
Styles | scabrous. |
hirtellous. |
Corollas | yellow, with scattered red spots, palate ridges with 2 white patches at tips, bilaterally symmetric, strongly bilabiate; tube-throat funnelform, (12–)16–20(–24) mm, exserted beyond calyx margin; limb expanded 8–10 mm, lobes obovate-oblong, apex rounded to truncate. |
yellow, red-dotted within, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, (14–)16–24 mm, exserted (8–)10–15 mm beyond calyx margin; limb broadly expanded. |
Fruiting pedicels | 15–35 mm, densely glandular-villous, hairs 0.5–1.2(–1.5) mm, gland-tipped. |
10–35 mm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs. |
Fruiting calyces | plicate-angled, cylindric-urceolate, weakly inflated, 6–12 mm, densely glandular-villous, hairs 0.5–1.2(–1.5) mm, gland-tipped, lobes 1–2 mm, apex rounded to mucronate. |
straight-erect or nodding 45–100º, ovate-campanulate, inflated, sagittally compressed, 15–22(–25) mm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs, throat closing. |
Capsules | included, 5–9 mm. |
included, 8–12 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
= 28. |
Erythranthe jungermannioides |
Erythranthe grandis |
|
Phenology | Flowering May–Jul(–Aug). | Flowering (Apr–)May–Jul(–Sep). |
Habitat | Basalt crevices in seepage zones in vertical cliff faces and canyon walls. | Beaches, dunes, coastal bluffs, wet cliff faces, mud flats and seeps, marshes, drainage ditches, creeks, rarely in coastal sage scrub. |
Elevation | 100–400(–1200) m. (300–1300(–3900) ft.) | 0–200(–800) m. (0–700(–2600) ft.) |
Distribution |
OR; WA
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CA; OR
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Discussion | The occurrence of Erythranthe jungermannioides in the Columbia River Gorge of Klickitat County, Washington (the only record from the state of Washington), is based on an imprecise, unconfirmed observation from the early 1990s (Washington National Heritage Program 2005). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The densely, evenly puberulent vestiture of pedicels, calyces, and distal stems usually is diagnostic, especially in combination with the large flowers (corollas and mature calyces) and tall stature. Plants from scattered collections are much shorter than normal but have large corollas and characteristic vestiture. Erythranthe grandis characteristically occurs in coastal localities from southern California to northern Oregon but also is found in inland localities and habitats near the coast but well away from salt spray. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 396. | FNA vol. 17, p. 412. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus jungermannioides | Mimulus guttatus var. grandis, M. grandis, M. guttatus subsp. litoralis, M. langsdorffii var. grandis, M. procerus |
Name authority | (Suksdorf) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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