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Jungermann's monkeyflower, liverwort monkey-flower

bent-stem monkeyflower, Dudley's monkeyflower

Habit Perennials, stoloniferous, stolons thin, forming overwintering turions. Annuals, fibrous-rooted or filiform-taprooted.
Stems

decumbent to procumbent, simple or branching near base, 5–38(–60) cm, densely glandular-villous, hairs 0.5–1.2(–1.5) mm, gland-tipped, internodes evident.

ascending to decumbent or prostrate, geniculate at nodes, simple or diffusely branched, 5–60 cm, moderately villous, hairs 0.8–2 mm, multicellular, eglandular and also 0.1–0.3 mm, stipitate-glandular.

Leaves

cauline, basal not persistent;

petiole 2–5(–20) mm;

blade subpalmately to pinnately veined, broadly ovate to broadly lanceolate, 7–35(–40) × 8–25 mm, base rounded, margins sharply, irregularly dentate to denticulate, apex acute to obtuse, surfaces glandular-villous.

basal and cauline, basal usually deciduous by flowering;

petiole 2–10(–35) mm;

blade pinnately to subpinnately veined, broadly ovate or elliptic-ovate to triangular, 8–35 × 5–30 mm, base cuneate to rounded or subcordate, margins serrate or dentate, teeth 3–10 per side, apex acute to obtuse or rounded, surfaces moderately villous, hairs 0.8–2 mm, multicellular, eglandular, and 0.1–0.3 mm, stipitate-glandular.

Flowers

herkogamous, 2 or 3, from medial to distal nodes.

herkogamous, (1–)6–20, from all or medial to distal nodes.

Styles

scabrous.

glabrous.

Corollas

yellow, with scattered red spots, palate ridges with 2 white patches at tips, bilaterally symmetric, strongly bilabiate;

tube-throat funnelform, (12–)16–20(–24) mm, exserted beyond calyx margin;

limb expanded 8–10 mm, lobes obovate-oblong, apex rounded to truncate.

yellow, without white patches, throat red-spotted, spots concentrated or becoming coalescent into a somewhat discrete splotch at base of each of 3 abaxial lobes and sometimes 2 adaxial, bilaterally symmetric, ± bilabiate;

tube-throat cylindric, 9–12 mm, exserted beyond calyx margin;

limb expanded 10–18 mm diam.

Fruiting pedicels

15–35 mm, densely glandular-villous, hairs 0.5–1.2(–1.5) mm, gland-tipped.

12–26(–55) mm, moderately villous, hairs 0.8–2 mm, multicellular, eglandular and also 0.1–0.3 mm, stipitate-glandular.

Fruiting calyces

plicate-angled, cylindric-urceolate, weakly inflated, 6–12 mm, densely glandular-villous, hairs 0.5–1.2(–1.5) mm, gland-tipped, lobes 1–2 mm, apex rounded to mucronate.

red-spotted, campanulate-cylindric, weakly inflated, (5–)6–8 mm, margins distinctly toothed or lobed, sparsely to moderately villous-glandular, ribs shallowly wing-angled, lobes pronounced, erect to spreading or spreading-recurving.

Capsules

included, 5–9 mm.

included, 4–6(–7) mm.

Anthers

included, glabrous.

included, glabrous.

2n

= 32.

= 32.

Erythranthe jungermannioides

Erythranthe geniculata

Phenology Flowering May–Jul(–Aug). Flowering (Mar–)Apr–Jul.
Habitat Basalt crevices in seepage zones in vertical cliff faces and canyon walls. Granite crevices, canyon slopes, talus, crevices in volcanic outcrops, edges of boulders, roadsides, damp sandy soils, sandy water edges, gravelly soils and creek bottoms.
Elevation 100–400(–1200) m. (300–1300(–3900) ft.) 200–900(–1200) m. (700–3000(–3900) ft.)
Distribution
from FNA
OR; WA
[WildflowerSearch map]
from FNA
CA
[WildflowerSearch map]
Discussion

The occurrence of Erythranthe jungermannioides in the Columbia River Gorge of Klickitat County, Washington (the only record from the state of Washington), is based on an imprecise, unconfirmed observation from the early 1990s (Washington National Heritage Program 2005).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe geniculata is known from an apparently disjunct cluster of populations in Butte, Sutter, and Yuba counties and then from Tuolumne and Stanislaus counties south to Kern County.

Erythranthe geniculata, compared to E. floribunda, has larger, chasmogamous, and allogamous flowers. The anther pairs of E. geniculata are at different levels, and the stigma is slightly above the adaxial anther pair; in E. floribunda both anther pairs and the stigma are at the same level.

Erythranthe arenaria, E. geniculata, and E. norrisii constitute a group of apparently closely related species within sect. Mimulosma endemic along the Sierra Nevada. All have ovate-petiolate leaves (only the basal ones are sometimes ovate in E. arenaria) with pinnate to subpinnate venation. The more widespread E. floribunda, which is part of the above group, also is similar, but all three endemics have larger corollas with the tube exserted at greater length beyond the calyx margin.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 396. FNA vol. 17, p. 404.
Parent taxa Phrymaceae > Erythranthe Phrymaceae > Erythranthe
Sibling taxa
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms Mimulus jungermannioides Mimulus geniculatus, M. dudleyi, M. floribundus var. geniculatus
Name authority (Suksdorf) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) (Greene) G. L. Nesom: Phytoneuron 2012-39: 38. (2012)
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