Erythranthe jungermannioides |
Erythranthe brachystylis |
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Jungermann's monkeyflower, liverwort monkey-flower |
short-pedicel monkeyflower |
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Habit | Perennials, stoloniferous, stolons thin, forming overwintering turions. | Annuals, fibrous-rooted, sometimes taprooted, apparently sometimes producing thin runners from basal nodes. |
Stems | decumbent to procumbent, simple or branching near base, 5–38(–60) cm, densely glandular-villous, hairs 0.5–1.2(–1.5) mm, gland-tipped, internodes evident. |
erect, simple or branched from proximal to medial nodes, 4-angled, filiform to slightly thickened, not distinctly fistulose, 6–22 cm, glabrous. |
Leaves | cauline, basal not persistent; petiole 2–5(–20) mm; blade subpalmately to pinnately veined, broadly ovate to broadly lanceolate, 7–35(–40) × 8–25 mm, base rounded, margins sharply, irregularly dentate to denticulate, apex acute to obtuse, surfaces glandular-villous. |
basal and cauline; petiole: proximals 1–8 mm, distals 0 mm; blade palmately 3–5-veined, ovate to depressed-ovate or suborbicular, 10–40 × 6–25 mm, base truncate to subcordate, margins undulate, subentire, or weakly, irregularly dentate, apex rounded, surfaces: proximals and medials glabrous, distals villous, hairs thin-walled, flattened, vitreous and sharp-pointed, eglandular. |
Flowers | herkogamous, 2 or 3, from medial to distal nodes. |
plesiogamous, 4–10, from medial to distal nodes, cleistogamous. |
Styles | scabrous. |
glabrous. |
Corollas | yellow, with scattered red spots, palate ridges with 2 white patches at tips, bilaterally symmetric, strongly bilabiate; tube-throat funnelform, (12–)16–20(–24) mm, exserted beyond calyx margin; limb expanded 8–10 mm, lobes obovate-oblong, apex rounded to truncate. |
yellow, without red markings, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly cylindric, 7–9 mm, exserted 0–1 mm beyond calyx margin; limb expanded 3 mm. |
Fruiting pedicels | 15–35 mm, densely glandular-villous, hairs 0.5–1.2(–1.5) mm, gland-tipped. |
5–10 mm in proximal axils, shorter than or equal to subtending leaves, 1–5 mm distally and flowers and fruits appearing sessile or subsessile, glabrous. |
Fruiting calyces | plicate-angled, cylindric-urceolate, weakly inflated, 6–12 mm, densely glandular-villous, hairs 0.5–1.2(–1.5) mm, gland-tipped, lobes 1–2 mm, apex rounded to mucronate. |
red-tinged to sparsely purple-dotted or not, broadly elliptic-ovoid, inflated, sagittally compressed, 10–13 mm, minutely hirtellous, throat not or slightly closing. |
Capsules | included, 5–9 mm. |
included, stipitate, 4–5 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
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Erythranthe jungermannioides |
Erythranthe brachystylis |
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Phenology | Flowering May–Jul(–Aug). | Flowering Jun–Aug. |
Habitat | Basalt crevices in seepage zones in vertical cliff faces and canyon walls. | Around springs, steep slopes. |
Elevation | 100–400(–1200) m. (300–1300(–3900) ft.) | 2100 m. (6900 ft.) |
Distribution |
OR; WA
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NV |
Discussion | The occurrence of Erythranthe jungermannioides in the Columbia River Gorge of Klickitat County, Washington (the only record from the state of Washington), is based on an imprecise, unconfirmed observation from the early 1990s (Washington National Heritage Program 2005). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe brachystylis is closely similar to E. arvensis. Plants of both are annual in duration and produce depressed-ovate leaves, the distal with vitreous-villous surfaces, and tiny corollas barely exserted from the calyx and probably cleistogamous. Vestiture of the distal leaves includes an admixture of eglandular sharp-pointed hairs, sometimes encountered in E. arvensis, though not typical, perhaps reflecting introgression from E. nasuta. The distinction of Erythranthe brachystylis from E. arvensis is primarily in its foreshortened pedicels and more inflated fruiting calyces. The fruiting calyces appear to be subsessile or on pedicels shorter or only equaling the subtending leaves. The difference is essentially qualitative but produces a distinctive aspect. Erythranthe brachystylis is known only from the type collection in Nye County, a region where E. arvensis has not been documented. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 396. | FNA vol. 17, p. 422. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus jungermannioides | Mimulus brachystylis |
Name authority | (Suksdorf) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | (Edwin) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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