Erythranthe grayi |
Erythranthe scouleri |
|
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Gray's monkeyflower, mariposa monkeyflower |
Columbia River monkey-flower, Scouler's monkey-flower |
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Habit | Annuals, fibrous-rooted. | Perennials, rhizomatous, producing long, sparsely leafy runners from basal nodes. |
Stems | erect, simple or branched from base, weakly 4-angled, 8–20 cm, glabrous. |
erect, simple or few-branched, 15–80 cm, glabrous. |
Leaves | basal and cauline; petiole 0 mm; blade palmately 3–5-veined, broadly ovate, 7–18 × 5–12 mm, base rounded, margins denticulate, apex acute, surfaces glabrous, rarely sparsely puberulent. |
cauline, basal usually not persistent; petiole: proximals and medial 10–25 mm (gradually merging into blade), distals 0 mm; blade palmately (3–)5–7-veined to subpinnate, (proximal to medial) oblong-elliptic to oblong-lanceolate, 25–60 × 8–18 mm, usually 3–4 times longer than wide, base attenuate, margins evenly, shallowly dentate or crenate to mucronate or mucronulate, teeth 10–20 per side, sometimes more deeply toothed at base, apex obtuse to acute, surfaces glabrous. |
Flowers | herkogamous, 2–20, from proximal to distal nodes. |
herkogamous, (1 or)2–8, from distal nodes. |
Styles | glabrous. |
minutely, prominently hirsutulous to villosulous. |
Corollas | rose red, throat pink lined with rose red and a yellow patch, abaxial ridges yellow, bilaterally symmetric, weakly bilabiate; tube-throat cylindric-funnelform, 8–11 mm, exserted beyond calyx margin; limb expanded 7–10 mm. |
yellow, without red markings, bilaterally symmetric, bilabiate; tube-throat funnelform, 20–24 mm, exserted 10–15 mm beyond calyx margin; limb expanded 22–30 mm. |
Fruiting pedicels | 6–7 mm, shorter than subtending leaves, glabrous. |
20–25 mm, glabrous. |
Fruiting calyces | campanulate, 9–11 × 5–6 mm, margins subtruncate, glabrous, sometimes densely papillate at flowering with tiny, 1-celled, eglandular hairs, these apparently deciduous by fruiting, lobes reduced, subequal. |
ovoid, inflated, sagittally compressed, 13–14 mm, glabrous, throat closing. |
Capsules | included, 5–9 mm. |
unknown. |
Anthers | included, minutely villous-hirsute. |
included, glabrous. |
Erythranthe grayi |
Erythranthe scouleri |
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Phenology | Flowering May–Jul(–Oct). | Flowering May–Aug. |
Habitat | Drying pond beds, creek banks, yellow pine, yellow pine-Libocedrus woodlands. | Salt marshes, river banks. |
Elevation | 1000–1900 m. (3300–6200 ft.) | 0–20 m. (0–100 ft.) |
Distribution |
CA
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OR; WA |
Discussion | In addition to the features noted in the key and descriptions, the fruiting calyces of Erythranthe grayi are distinctly more inflated than those of E. acutidens and E. inconspicua. G. L. Nesom (2012g) maintained E. grayi as distinct from E. acutidens, relying primarily on fruiting pedicel length (see key above), but the two have nearly identical ranges (Tuolumne County south to Kern County), and study of additional collections suggests that only a single species may be represented. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe scouleri is distinctive in its oblong-elliptic leaves with long-tapering bases and closely toothed margins, completely glabrous vestiture, long, leafy runners from basal cauline nodes, large corollas with broad limbs, and prominently hairy styles. Several features suggest a close relationship to E. decora, particularly its very large corollas, hairy styles, closely toothed leaf margins, tall, simple, and erect stems, numerous runners, and its geographic range. All collections apparently have been made near the mouth of the Columbia River in Clatsop and Columbia counties, Oregon (G. L. Nesom 2013d). Recent observations (Alexander John Wright, pers. comm.; photos http://www.inaturalist.org/observations/3537008) indicate that it also occurs in Wahkiakum County, Washington, in the Julia Butler Hansen Refuge. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 382. | FNA vol. 17, p. 415. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus grayi | Mimulus scouleri, M. guttatus subsp. scouleri |
Name authority | (A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 34. (2012) | (Hooker) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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