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large monkey-flower, magnificent monkeyflower, magnificent seep monkeyflower

fern-leaf monkeyflower

Habit Perennials, rhizomatous, sometimes rooting at proximal nodes. Annuals, fibrous-rooted.
Stems

erect, sometimes decumbent basally, branched, often fistulose, (25–)50–120(–160) cm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs.

erect or slightly ascending from base, sometimes purplish, simple or few-branched from proximal nodes, 3–38 cm, glabrous.

Leaves

basal and cauline, basal usually not persistent, bracteate in inflorescence;

petiole 10–80 mm, gradually reduced distally;

blade subpinnately, sometimes palmately, 5–7-veined, ovate to broadly elliptic, 25–60 × 20–40(–60) mm, usually 1–2 times longer than wide, base truncate or truncate-cuneate to subcordate, margins crenulate to dentate, proximally sometimes sublyrate, apex rounded to obtuse, surfaces of distals densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs.

basal and cauline;

petiole (proximal and proximal to mid cauline), distal bracteate or absent, ovate, margins entire;

blade: divisions 1-veined, oblong-lanceolate to ovate in outline, 3–68 mm, slightly thickened, base (at proximalmost division) truncate, margins (of divisions) entire, bipinnately dissected, ca. 5–12(–15) primary divisions on each side, ultimate division nearly linear, apex obtuse to acute, surfaces glabrous.

Flowers

herkogamous, 8–26, mostly from distal nodes, usually in bracteate racemes.

plesiogamous, 4–10(–12), on foreshortened distal nodes, usually clustered, chasmogamous, sometimes cleistogamous.

Styles

hirtellous.

glabrous.

Corollas

yellow, red-dotted within, bilaterally symmetric, bilabiate;

tube-throat broadly funnelform, (14–)16–24 mm, exserted (8–)10–15 mm beyond calyx margin;

limb broadly expanded.

yellow, proximal lip sometimes red-splotched, palate ridges yellow, bilaterally symmetric, bilabiate;

tube-throat cylindric-funnelform, 4–8 mm, exserted 0–2 mm beyond calyx margin or not; palate ridges densely villous.

Fruiting pedicels

10–35 mm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs.

6–14 mm, glabrous.

Fruiting calyces

straight-erect or nodding 45–100º, ovate-campanulate, inflated, sagittally compressed, 15–22(–25) mm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs, throat closing.

urceolate to urceolate-campanulate, inflated, sagittally compressed, 8–11 mm, glabrous, throat closing.

Capsules

included, 8–12 mm.

included, 3–8 mm.

Anthers

included, glabrous.

included, glabrous.

2n

= 28.

Erythranthe grandis

Erythranthe filicifolia

Phenology Flowering (Apr–)May–Jul(–Sep). Flowering Apr–Jun(–Sep).
Habitat Beaches, dunes, coastal bluffs, wet cliff faces, mud flats and seeps, marshes, drainage ditches, creeks, rarely in coastal sage scrub. Slow-draining, ephemeral seeps on exfoliating granite slabs, over basalt.
Elevation 0–200(–800) m. (0–700(–2600) ft.) 300–1600 m. (1000–5200 ft.)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
from FNA
CA
Discussion

The densely, evenly puberulent vestiture of pedicels, calyces, and distal stems usually is diagnostic, especially in combination with the large flowers (corollas and mature calyces) and tall stature. Plants from scattered collections are much shorter than normal but have large corollas and characteristic vestiture.

Erythranthe grandis characteristically occurs in coastal localities from southern California to northern Oregon but also is found in inland localities and habitats near the coast but well away from salt spray.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe filicifolia is known only from localities in Butte and Plumas counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 412. FNA vol. 17, p. 418.
Parent taxa Phrymaceae > Erythranthe Phrymaceae > Erythranthe
Sibling taxa
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms Mimulus guttatus var. grandis, M. grandis, M. guttatus subsp. litoralis, M. langsdorffii var. grandis, M. procerus Mimulus filicifolius
Name authority (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) (Sexton: K. G. Ferris & Schoenig) G. L. Nesom, Phytoneuron 2013-80: 1. (2013)
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