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large monkey-flower, magnificent monkeyflower, magnificent seep monkeyflower

Palomar monkeyflower

Habit Perennials, rhizomatous, sometimes rooting at proximal nodes. Annuals, taprooted.
Stems

erect, sometimes decumbent basally, branched, often fistulose, (25–)50–120(–160) cm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs.

erect, simple or branched from basal nodes, 4–20 cm, minutely puberulent.

Leaves

basal and cauline, basal usually not persistent, bracteate in inflorescence;

petiole 10–80 mm, gradually reduced distally;

blade subpinnately, sometimes palmately, 5–7-veined, ovate to broadly elliptic, 25–60 × 20–40(–60) mm, usually 1–2 times longer than wide, base truncate or truncate-cuneate to subcordate, margins crenulate to dentate, proximally sometimes sublyrate, apex rounded to obtuse, surfaces of distals densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs.

cauline, basal not persistent;

petiole 0 mm;

blade 1-veined or palmately 3-veined (in broader ones), lanceolate to ovate, 3–20 × 1–6 mm, base attenuate, margins entire, sometimes toothed, apex acute, surfaces minutely puberulent.

Flowers

herkogamous, 8–26, mostly from distal nodes, usually in bracteate racemes.

herkogamous, 1–36, from distal or medial to distal nodes.

Styles

hirtellous.

distally pubescent.

Corollas

yellow, red-dotted within, bilaterally symmetric, bilabiate;

tube-throat broadly funnelform, (14–)16–24 mm, exserted (8–)10–15 mm beyond calyx margin;

limb broadly expanded.

pink to purple, abaxial limb with 2 yellow palate ridges, bilaterally symmetric, weakly bilabiate;

tube-throat funnelform, (3–)6–10 mm, exserted beyond calyx margin;

limb expanded 3–14 mm, lobes notched, adaxial limb glabrous, sometimes sparsely bearded.

Fruiting pedicels

10–35 mm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs.

ascending to spreading horizontally, (2–)12–60(–68) mm.

Fruiting calyces

straight-erect or nodding 45–100º, ovate-campanulate, inflated, sagittally compressed, 15–22(–25) mm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs, throat closing.

sometimes red-dotted, campanulate, 3–6 mm, margins distinctly toothed or lobed, minutely puberulent, ribs weak, lobes pronounced, erect, margins glabrous.

Capsules

included, 8–12 mm.

included, 5–8 mm.

Anthers

included, glabrous.

included, glabrous.

Stigmas

distinctly shorter than corolla tube.

2n

= 28.

Erythranthe grandis

Erythranthe diffusa

Phenology Flowering (Apr–)May–Jul(–Sep). Flowering Apr–Jun.
Habitat Beaches, dunes, coastal bluffs, wet cliff faces, mud flats and seeps, marshes, drainage ditches, creeks, rarely in coastal sage scrub. Moist areas in openings of chaparral, dry meadows in pine and oak woodlands.
Elevation 0–200(–800) m. (0–700(–2600) ft.) 300–1800 m. (1000–5900 ft.)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
Discussion

The densely, evenly puberulent vestiture of pedicels, calyces, and distal stems usually is diagnostic, especially in combination with the large flowers (corollas and mature calyces) and tall stature. Plants from scattered collections are much shorter than normal but have large corollas and characteristic vestiture.

Erythranthe grandis characteristically occurs in coastal localities from southern California to northern Oregon but also is found in inland localities and habitats near the coast but well away from salt spray.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe diffusa has been included previously in E. palmeri but can be distinguished morphologically by glabrous margins on the calyx and pubescence on the distal end of the style. It occurs in Orange, Riverside, and San Diego counties as well as close to the California border in Baja California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 412. FNA vol. 17, p. 387.
Parent taxa Phrymaceae > Erythranthe Phrymaceae > Erythranthe
Sibling taxa
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms Mimulus guttatus var. grandis, M. grandis, M. guttatus subsp. litoralis, M. langsdorffii var. grandis, M. procerus Mimulus diffusus, M. grantianus
Name authority (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) (A. L. Grant) N. S. Fraga: Phytoneuron 2012-39: 35. (2012)
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