Erythranthe glaucescens |
Erythranthe thermalis |
|
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shield-bract monkeyflower |
Yellowstone monkeyflower |
|
Habit | Annuals, slender-taprooted or fibrous-rooted, rarely with runners from basal nodes. | Annuals, taprooted, rarely with a basal, runnerlike stem. |
Stems | erect, simple or branched, terete, sometimes 4-angled distally, (5–)30–60(–80) cm, glabrous, glaucous. |
erect, simple or branched from basal nodes, 1.5–10(–15) cm, villous-glandular proximally, moderately to densely stipitate-glandular, sometimes hirtellous and eglandular distally. |
Leaves | basal and cauline; petiole: basal and proximal cauline as long as or much longer than blade, slender, sometimes pubescent or villous, distals absent; blade palmately 3–5-veined, (proximal) ovate to ovate-elliptic or orbicular-ovate, sometimes subcordate, 10–50 mm, midcauline to distal orbicular, 5–45 mm wide, distinctly connate-perfoliate, disclike distally, base rounded to subcordate, margins: proximals denticulate to dentate or coarsely, irregularly toothed, sometimes lobed at base, distals nearly entire or toothed, teeth scattered, small, apex rounded, surfaces glabrous, glaucous. |
basal and cauline, cauline 2–5 pairs; petiole: basal and proximal cauline 3–20 mm, distals 0 mm; blade palmately 3–5-veined, suborbicular to ovate, depressed-ovate, ovate-deltate, or reniform, 4–15(–20) × 4–20 mm, base cuneate to truncate or subcordate, margins evenly crenate-dentate to subentire, apex acute to obtuse or rounded, surfaces moderately to densely stipitate-glandular, sometimes hirtellous and eglandular. |
Flowers | herkogamous, 1–16, from distal nodes, sometimes from nearly all, chasmogamous. |
plesiogamous, 1–5(–9), usually at distal nodes, chasmogamous. |
Styles | minutely hirtellous-puberulent. |
hirtellous. |
Corollas | yellow, sometimes with a median splotch, abaxial limb densely dark yellow, others much lighter, throat floor and tube red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 12–23 mm, exserted 4–8 mm beyond calyx margin; limb expanded 14–36 mm. |
yellow, red-dotted or not, bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–12 mm, exserted 1–2 mm beyond calyx margin; limb expanded 12–15 mm, throat open, palate villous. |
Fruiting pedicels | 10–50 mm, glabrous, glaucous. |
7–12 mm, moderately to densely stipitate-glandular, sometimes hirtellous and eglandular. |
Fruiting calyces | broadly campanulate, inflated, sagittally compressed, 7–16 mm, glabrous, glaucous, throat closing. |
ovate-campanulate, inflated, sagittally compressed, 8–11 mm, moderately to densely stipitate-glandular, sometimes hirtellous and eglandular, throat closing, adaxial lobe longer than others. |
Capsules | included, 5–11 mm. |
included, 5–6 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28. |
= 28. |
Erythranthe glaucescens |
Erythranthe thermalis |
|
Phenology | Flowering Mar–May(–Jun). | Flowering Mar–Aug. |
Habitat | Seepage areas, wet rocks, moist cliffs, pool edges, gravelly stream banks, serpentine outcrops, roadsides and roadcuts, low pastures, riparian woodlands, blue oak woodlands, chaparral, grasslands. | Hot, shallow, quick-drying soils around thermal pools and vents. |
Elevation | 80–900(–1100) m. (300–3000(–3600) ft.) | 2200–2600 m. (7200–8500 ft.) |
Distribution |
CA
|
WY |
Discussion | Plants from one locality in Butte County are unusual in producing filiform, small-leaved runners from basal cauline nodes. Erythranthe glaucescens is known only from Butte and Tehama counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe thermalis is endemic to Yellowstone National Park in northwestern Wyoming. The species is recognized by its annual duration (without rhizomes), reduced stature and leaf size, and short, but broad-limbed, corollas with autogamous fertilization. Typical E. guttata (rhizomatous, herkogamous) also grows in the immediately surrounding areas but apparently not in hot soils. Each species maintains distinctions in growth form, phenology and mating system in common garden experiments (Y. Lekberg et al. 2012). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 417. | FNA vol. 17, p. 413. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus glaucescens, M. guttatus var. glaucescens | Mimulus thermalis |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) | (A. Nelson) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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