FNA: Erythranthe glaucescens vs. Erythranthe sierrae

	Erythranthe glaucescens	Erythranthe sierrae
	shield-bract monkeyflower	Sierra Nevada monkeyflower
Habit	Annuals, slender-taprooted or fibrous-rooted, rarely with runners from basal nodes.	Annuals, taprooted.
Stems	erect, simple or branched, terete, sometimes 4-angled distally, (5–)30–60(–80) cm, glabrous, glaucous.	erect, simple or branched from basal nodes, (3–)4–20 cm, minutely glandular.
Leaves	basal and cauline; petiole: basal and proximal cauline as long as or much longer than blade, slender, sometimes pubescent or villous, distals absent; blade palmately 3–5-veined, (proximal) ovate to ovate-elliptic or orbicular-ovate, sometimes subcordate, 10–50 mm, midcauline to distal orbicular, 5–45 mm wide, distinctly connate-perfoliate, disclike distally, base rounded to subcordate, margins: proximals denticulate to dentate or coarsely, irregularly toothed, sometimes lobed at base, distals nearly entire or toothed, teeth scattered, small, apex rounded, surfaces glabrous, glaucous.	cauline, basal not persistent; petiole 0 mm; blade 1-veined or palmately 3-veined (in broader ones), linear to oblanceolate, 3–23 × 1–10 mm, base attenuate, margins entire, sometimes toothed, apex acute, surfaces minutely glandular.
Flowers	herkogamous, 1–16, from distal nodes, sometimes from nearly all, chasmogamous.	herkogamous, 1–38, from distal or medial to distal nodes.
Styles	minutely hirtellous-puberulent.	glabrous.
Corollas	yellow, sometimes with a median splotch, abaxial limb densely dark yellow, others much lighter, throat floor and tube red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 12–23 mm, exserted 4–8 mm beyond calyx margin; limb expanded 14–36 mm.	pale pink, abaxial limb with 2 yellow ridges, bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 5–15 mm, exserted beyond calyx margin; limb expanded 5–17 mm, lobes deeply notched, abaxial limb glabrous or sparsely puberulent.
Fruiting pedicels	10–50 mm, glabrous, glaucous.	10–33(–40) mm.
Fruiting calyces	broadly campanulate, inflated, sagittally compressed, 7–16 mm, glabrous, glaucous, throat closing.	sometimes red-spotted, campanulate to cylindric, 4–8 mm, margins distinctly toothed or lobed, minutely glandular, ribs weak, lobes pronounced, erect, margins ciliate.
Capsules	included, 5–11 mm.	included, 4–9 mm.
Anthers	included, glabrous.	included, glabrous.
2n	= 28.

	Erythranthe glaucescens	Erythranthe sierrae
Phenology	Flowering Mar–May(–Jun).	Flowering Apr–Jul.
Habitat	Seepage areas, wet rocks, moist cliffs, pool edges, gravelly stream banks, serpentine outcrops, roadsides and roadcuts, low pastures, riparian woodlands, blue oak woodlands, chaparral, grasslands.	Foothill oak woodlands, mixed coniferous forests.
Elevation	80–900(–1100) m. (300–3000(–3600) ft.)	200–2300 m. (700–7500 ft.)
Distribution	CA [WildflowerSearch map]	CA
Discussion	Plants from one locality in Butte County are unusual in producing filiform, small-leaved runners from basal cauline nodes. Erythranthe glaucescens is known only from Butte and Tehama counties. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Erythranthe sierrae is endemic to the southern Sierra Nevada in Fresno, Kern, and Tulare counties. The species was previously treated as E. palmeri but can be distinguished by having leaf margins that are often toothed, pale pink corollas, and white stamens. Erythranthe palmeri has entire leaf margins that are never toothed, deep pink to purple corollas, and yellow stamens. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 417.	FNA vol. 17, p. 385.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus glaucescens, M. guttatus var. glaucescens
Name authority	(Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)	N. S. Fraga: Aliso 30: 67, figs. 29–31. (2012)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe sierrae

shield-bract monkeyflower

Sierra Nevada monkeyflower

Habit

Annuals, slender-taprooted or fibrous-rooted, rarely with runners from basal nodes.

Annuals, taprooted.

Stems

erect, simple or branched, terete, sometimes 4-angled distally, (5–)30–60(–80) cm, glabrous, glaucous.

erect, simple or branched from basal nodes, (3–)4–20 cm, minutely glandular.

Leaves

basal and cauline;

petiole: basal and proximal cauline as long as or much longer than blade, slender, sometimes pubescent or villous, distals absent;

blade palmately 3–5-veined, (proximal) ovate to ovate-elliptic or orbicular-ovate, sometimes subcordate, 10–50 mm, midcauline to distal orbicular, 5–45 mm wide, distinctly connate-perfoliate, disclike distally, base rounded to subcordate, margins: proximals denticulate to dentate or coarsely, irregularly toothed, sometimes lobed at base, distals nearly entire or toothed, teeth scattered, small, apex rounded, surfaces glabrous, glaucous.

cauline, basal not persistent;

petiole 0 mm;

blade 1-veined or palmately 3-veined (in broader ones), linear to oblanceolate, 3–23 × 1–10 mm, base attenuate, margins entire, sometimes toothed, apex acute, surfaces minutely glandular.

Flowers

herkogamous, 1–16, from distal nodes, sometimes from nearly all, chasmogamous.

herkogamous, 1–38, from distal or medial to distal nodes.

Styles

minutely hirtellous-puberulent.

glabrous.

Corollas

yellow, sometimes with a median splotch, abaxial limb densely dark yellow, others much lighter, throat floor and tube red-dotted, bilaterally symmetric, bilabiate;

tube-throat funnelform, 12–23 mm, exserted 4–8 mm beyond calyx margin;

limb expanded 14–36 mm.

pale pink, abaxial limb with 2 yellow ridges, bilaterally symmetric, strongly bilabiate;

tube-throat funnelform, 5–15 mm, exserted beyond calyx margin;

limb expanded 5–17 mm, lobes deeply notched, abaxial limb glabrous or sparsely puberulent.

Fruiting pedicels

10–50 mm, glabrous, glaucous.

10–33(–40) mm.

Fruiting calyces

broadly campanulate, inflated, sagittally compressed, 7–16 mm, glabrous, glaucous, throat closing.

sometimes red-spotted, campanulate to cylindric, 4–8 mm, margins distinctly toothed or lobed, minutely glandular, ribs weak, lobes pronounced, erect, margins ciliate.

Capsules

included, 5–11 mm.

included, 4–9 mm.

Anthers

included, glabrous.

= 28.

Erythranthe glaucescens

Erythranthe sierrae

Phenology

Flowering Mar–May(–Jun).

Flowering Apr–Jul.

Habitat

Seepage areas, wet rocks, moist cliffs, pool edges, gravelly stream banks, serpentine outcrops, roadsides and roadcuts, low pastures, riparian woodlands, blue oak woodlands, chaparral, grasslands.

Foothill oak woodlands, mixed coniferous forests.

Elevation

80–900(–1100) m. (300–3000(–3600) ft.)

200–2300 m. (700–7500 ft.)

Distribution

[WildflowerSearch map]

Discussion

Plants from one locality in Butte County are unusual in producing filiform, small-leaved runners from basal cauline nodes. Erythranthe glaucescens is known only from Butte and Tehama counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe sierrae is endemic to the southern Sierra Nevada in Fresno, Kern, and Tulare counties. The species was previously treated as E. palmeri but can be distinguished by having leaf margins that are often toothed, pale pink corollas, and white stamens. Erythranthe palmeri has entire leaf margins that are never toothed, deep pink to purple corollas, and yellow stamens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 417.

FNA vol. 17, p. 385.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus glaucescens, M. guttatus var. glaucescens

Name authority

(Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)

N. S. Fraga: Aliso 30: 67, figs. 29–31. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Erythranthe glaucescens

Erythranthe sierrae

Erythranthe glaucescens

Erythranthe sierrae