Erythranthe glaucescens |
Erythranthe regni |
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shield-bract monkeyflower |
King of Arizona monkeyflower |
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Habit | Annuals, slender-taprooted or fibrous-rooted, rarely with runners from basal nodes. | Annuals, fibrous-rooted, sometimes rooting at proximal nodes. |
Stems | erect, simple or branched, terete, sometimes 4-angled distally, (5–)30–60(–80) cm, glabrous, glaucous. |
erect to ascending-erect, branched, sometimes becoming slightly fistulose, 15–45 cm, glabrous. |
Leaves | basal and cauline; petiole: basal and proximal cauline as long as or much longer than blade, slender, sometimes pubescent or villous, distals absent; blade palmately 3–5-veined, (proximal) ovate to ovate-elliptic or orbicular-ovate, sometimes subcordate, 10–50 mm, midcauline to distal orbicular, 5–45 mm wide, distinctly connate-perfoliate, disclike distally, base rounded to subcordate, margins: proximals denticulate to dentate or coarsely, irregularly toothed, sometimes lobed at base, distals nearly entire or toothed, teeth scattered, small, apex rounded, surfaces glabrous, glaucous. |
basal and cauline; petiole: proximals 5–25(–30) mm, mid cauline and distals not connate, 0 mm; blade palmately 5–7-veined, proximal sometimes subpinnate, proximals ovate to depressed-orbicular, 15–20(–50) × 15–25(–50) mm, medials and distals broadly depressed-ovate to obtriangular or flabellate, 15–35 mm, largest basal or at mid stem with distal slightly reduced, base attenuate-cuneate, margins shallowly serrate-dentate, sometimes irregularly, to mucronulate or apiculate, teeth (3–)5–7 per side, rarely subentire, apex rounded, surfaces glabrous. |
Flowers | herkogamous, 1–16, from distal nodes, sometimes from nearly all, chasmogamous. |
plesiogamous, 6–16, from all nodes or medial to distal, cleistogamous. |
Styles | minutely hirtellous-puberulent. |
glabrous. |
Corollas | yellow, sometimes with a median splotch, abaxial limb densely dark yellow, others much lighter, throat floor and tube red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 12–23 mm, exserted 4–8 mm beyond calyx margin; limb expanded 14–36 mm. |
yellow, not red-dotted, bilaterally or radially symmetric, bilabiate or regular; tube-throat cylindric-funnelform, 9–12 mm, exserted 3–5 mm beyond calyx margin; limb expanded 1–1.5 mm. |
Fruiting pedicels | 10–50 mm, glabrous, glaucous. |
15–30 mm, longer than subtending leaves, glabrous. |
Fruiting calyces | broadly campanulate, inflated, sagittally compressed, 7–16 mm, glabrous, glaucous, throat closing. |
sparsely purple-dotted, broadly campanulate-cylindric, inflated, sagittally compressed, 7–9 mm, glabrous, throat not closing, adaxial lobe longest. |
Capsules | included, 5–11 mm. |
included, 4–5 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28. |
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Erythranthe glaucescens |
Erythranthe regni |
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Phenology | Flowering Mar–May(–Jun). | Flowering Mar–May. |
Habitat | Seepage areas, wet rocks, moist cliffs, pool edges, gravelly stream banks, serpentine outcrops, roadsides and roadcuts, low pastures, riparian woodlands, blue oak woodlands, chaparral, grasslands. | Moist to wet, sandy loam soils. |
Elevation | 80–900(–1100) m. (300–3000(–3600) ft.) | 800–1000 m. (2600–3300 ft.) |
Distribution |
CA
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AZ |
Discussion | Plants from one locality in Butte County are unusual in producing filiform, small-leaved runners from basal cauline nodes. Erythranthe glaucescens is known only from Butte and Tehama counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe regni is endemic to the Kofa Mountains of Yuma County; all collections have been made from the Kofa Game Refuge (Kofa National Wildlife Refuge). Because its calyces remain open at maturity, this species is hypothesized to be most closely related to E. geyeri, from which it differs by its erect habit, apparently annual duration, larger leaves, purple-dotted calyces, and corollas with longer tube-throat and barely bilabiate limb. Geography and other morphology, however, suggest that its evolutionary origins are closer to E. guttata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 417. | FNA vol. 17, p. 406. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus glaucescens, M. guttatus var. glaucescens | |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) | G. L. Nesom: Phytoneuron 2012-40: 24. (2012) |
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