FNA: Erythranthe glaucescens vs. Erythranthe pardalis

	Erythranthe glaucescens	Erythranthe pardalis
	shield-bract monkeyflower	Pennell's panther
Habit	Annuals, slender-taprooted or fibrous-rooted, rarely with runners from basal nodes.	Annuals, fibrous-rooted or taprooted.
Stems	erect, simple or branched, terete, sometimes 4-angled distally, (5–)30–60(–80) cm, glabrous, glaucous.	decumbent-ascending, erect distally, simple, sometimes branched from proximal to medial nodes, 5–30 cm, short, delicately stipitate-glandular, distals minutely puberulent-glandular, hairs 0.1–0.4 mm (to 1 mm on proximal portions of stems), gland-tipped.
Leaves	basal and cauline; petiole: basal and proximal cauline as long as or much longer than blade, slender, sometimes pubescent or villous, distals absent; blade palmately 3–5-veined, (proximal) ovate to ovate-elliptic or orbicular-ovate, sometimes subcordate, 10–50 mm, midcauline to distal orbicular, 5–45 mm wide, distinctly connate-perfoliate, disclike distally, base rounded to subcordate, margins: proximals denticulate to dentate or coarsely, irregularly toothed, sometimes lobed at base, distals nearly entire or toothed, teeth scattered, small, apex rounded, surfaces glabrous, glaucous.	usually cauline, basal usually not persistent; petiole: proximals and medials 8–20 mm, distalmost 1–2 mm; blade palmately 3-veined, usually ovate or broadly ovate to depressed-ovate, proximals and medials 7–22 × 6–18 mm, sometimes largest at mid stem, base rounded or cuneate to gradually attenuate, margins shallowly dentate-serrate, teeth 2 or 3(–5) per side mostly distally, apex obtuse to obtuse-acuminate, surfaces sparsely villous to puberulent-glandular, hairs vitreous, gland-tipped, sometimes glabrous.
Flowers	herkogamous, 1–16, from distal nodes, sometimes from nearly all, chasmogamous.	plesiogamous, 2–12, usually evenly distributed from proximal to distal nodes, chasmogamous, anther pairs in larger corollas slightly separated, stigma at level of distal pair, or both anther pairs and stigma at same level; in smaller corollas without expanded limb and barely exserted beyond calyx margin, both anther pairs and stigma at same level.
Styles	minutely hirtellous-puberulent.	glabrous.
Corollas	yellow, sometimes with a median splotch, abaxial limb densely dark yellow, others much lighter, throat floor and tube red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 12–23 mm, exserted 4–8 mm beyond calyx margin; limb expanded 14–36 mm.	yellow, throat floor sometimes red-spotted, bilaterally symmetric, bilabiate; tube-throat narrowly funnelform to cylindric, 7–10(–12) mm, exserted 1–3 mm beyond calyx margin; limb expanded 8–12 mm, palate villous.
Fruiting pedicels	10–50 mm, glabrous, glaucous.	10–35 mm, short, delicately stipitate-glandular, distals minutely puberulent-glandular, hairs 0.1–0.4 mm, gland-tipped.
Fruiting calyces	broadly campanulate, inflated, sagittally compressed, 7–16 mm, glabrous, glaucous, throat closing.	nodding 45–180º, consistently dark purple-spotted, cylindric-campanulate, inflated, sagittally compressed, 8–11 mm, glabrous or sparsely puberulent-glandular, sometimes minutely hirtellous, throat closing.
Capsules	included, 5–11 mm.	included, stipitate, 4–6 mm.
Anthers	included, glabrous.	included, glabrous.
2n	= 28.	= 28.

	Erythranthe glaucescens	Erythranthe pardalis
Phenology	Flowering Mar–May(–Jun).	Flowering (Mar–)Apr–May.
Habitat	Seepage areas, wet rocks, moist cliffs, pool edges, gravelly stream banks, serpentine outcrops, roadsides and roadcuts, low pastures, riparian woodlands, blue oak woodlands, chaparral, grasslands.	Crevices of serpentine rock, stony red soils, red clay, among boulders, along streams, ditches, tailings at copper mines.
Elevation	80–900(–1100) m. (300–3000(–3600) ft.)	100–700 m. (300–2300 ft.)
Distribution	CA [WildflowerSearch map]	CA
Discussion	Plants from one locality in Butte County are unusual in producing filiform, small-leaved runners from basal cauline nodes. Erythranthe glaucescens is known only from Butte and Tehama counties. (Discussion copyrighted by Flora of North America; reprinted with permission.)	The relative constancy of Erythranthe pardalis in morphology suggests that genetic influence from other species is slight. It is recognized by its annual duration and relatively delicate habit, ovate to depressed-ovate leaves toothed mostly on the distal margins, small flowers produced from all nodes (proximal to distal), dark-spotted calyces, and stipitate-glandular cauline and foliar vestiture. While the corolla limbs are distinctly expanded, the tubes are only slightly exserted from the calyx, and flowers apparently are plesiogamous. The epithet pardalis alludes to the dark-spotted calyx. Plants of Erythranthe pardalis occur primarily on serpentine rocks and soils but also grow on copper tailings at mine sites. The species is known from Amador, Calaveras, El Dorado, Placer, Tehama, and Tuolumne counties. The plants in Tehama County, geographically and ecologically disjunct from the main range, were recorded as growing in basalt crevices. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 417.	FNA vol. 17, p. 420.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus glaucescens, M. guttatus var. glaucescens	Mimulus pardalis, M. cupriphilus, M. guttatus var. cupriphilus, M. guttatus var. pardalis
Name authority	(Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)	(Pennell) G. L. Nesom: Phytoneuron 2012-39: 44. (2012)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe pardalis

shield-bract monkeyflower

Pennell's panther

Habit

Annuals, slender-taprooted or fibrous-rooted, rarely with runners from basal nodes.

Annuals, fibrous-rooted or taprooted.

Stems

erect, simple or branched, terete, sometimes 4-angled distally, (5–)30–60(–80) cm, glabrous, glaucous.

decumbent-ascending, erect distally, simple, sometimes branched from proximal to medial nodes, 5–30 cm, short, delicately stipitate-glandular, distals minutely puberulent-glandular, hairs 0.1–0.4 mm (to 1 mm on proximal portions of stems), gland-tipped.

Leaves

basal and cauline;

petiole: basal and proximal cauline as long as or much longer than blade, slender, sometimes pubescent or villous, distals absent;

blade palmately 3–5-veined, (proximal) ovate to ovate-elliptic or orbicular-ovate, sometimes subcordate, 10–50 mm, midcauline to distal orbicular, 5–45 mm wide, distinctly connate-perfoliate, disclike distally, base rounded to subcordate, margins: proximals denticulate to dentate or coarsely, irregularly toothed, sometimes lobed at base, distals nearly entire or toothed, teeth scattered, small, apex rounded, surfaces glabrous, glaucous.

usually cauline, basal usually not persistent;

petiole: proximals and medials 8–20 mm, distalmost 1–2 mm;

blade palmately 3-veined, usually ovate or broadly ovate to depressed-ovate, proximals and medials 7–22 × 6–18 mm, sometimes largest at mid stem, base rounded or cuneate to gradually attenuate, margins shallowly dentate-serrate, teeth 2 or 3(–5) per side mostly distally, apex obtuse to obtuse-acuminate, surfaces sparsely villous to puberulent-glandular, hairs vitreous, gland-tipped, sometimes glabrous.

Flowers

herkogamous, 1–16, from distal nodes, sometimes from nearly all, chasmogamous.

plesiogamous, 2–12, usually evenly distributed from proximal to distal nodes, chasmogamous, anther pairs in larger corollas slightly separated, stigma at level of distal pair, or both anther pairs and stigma at same level; in smaller corollas without expanded limb and barely exserted beyond calyx margin, both anther pairs and stigma at same level.

Styles

minutely hirtellous-puberulent.

glabrous.

Corollas

yellow, sometimes with a median splotch, abaxial limb densely dark yellow, others much lighter, throat floor and tube red-dotted, bilaterally symmetric, bilabiate;

tube-throat funnelform, 12–23 mm, exserted 4–8 mm beyond calyx margin;

limb expanded 14–36 mm.

yellow, throat floor sometimes red-spotted, bilaterally symmetric, bilabiate;

tube-throat narrowly funnelform to cylindric, 7–10(–12) mm, exserted 1–3 mm beyond calyx margin;

limb expanded 8–12 mm, palate villous.

Fruiting pedicels

10–50 mm, glabrous, glaucous.

10–35 mm, short, delicately stipitate-glandular, distals minutely puberulent-glandular, hairs 0.1–0.4 mm, gland-tipped.

Fruiting calyces

broadly campanulate, inflated, sagittally compressed, 7–16 mm, glabrous, glaucous, throat closing.

nodding 45–180º, consistently dark purple-spotted, cylindric-campanulate, inflated, sagittally compressed, 8–11 mm, glabrous or sparsely puberulent-glandular, sometimes minutely hirtellous, throat closing.

Capsules

included, 5–11 mm.

included, stipitate, 4–6 mm.

Anthers

included, glabrous.

= 28.

Erythranthe glaucescens

Erythranthe pardalis

Phenology

Flowering Mar–May(–Jun).

Flowering (Mar–)Apr–May.

Habitat

Seepage areas, wet rocks, moist cliffs, pool edges, gravelly stream banks, serpentine outcrops, roadsides and roadcuts, low pastures, riparian woodlands, blue oak woodlands, chaparral, grasslands.

Crevices of serpentine rock, stony red soils, red clay, among boulders, along streams, ditches, tailings at copper mines.

Elevation

80–900(–1100) m. (300–3000(–3600) ft.)

100–700 m. (300–2300 ft.)

Distribution

[WildflowerSearch map]

Discussion

Plants from one locality in Butte County are unusual in producing filiform, small-leaved runners from basal cauline nodes. Erythranthe glaucescens is known only from Butte and Tehama counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The relative constancy of Erythranthe pardalis in morphology suggests that genetic influence from other species is slight. It is recognized by its annual duration and relatively delicate habit, ovate to depressed-ovate leaves toothed mostly on the distal margins, small flowers produced from all nodes (proximal to distal), dark-spotted calyces, and stipitate-glandular cauline and foliar vestiture. While the corolla limbs are distinctly expanded, the tubes are only slightly exserted from the calyx, and flowers apparently are plesiogamous. The epithet pardalis alludes to the dark-spotted calyx.

Plants of Erythranthe pardalis occur primarily on serpentine rocks and soils but also grow on copper tailings at mine sites. The species is known from Amador, Calaveras, El Dorado, Placer, Tehama, and Tuolumne counties. The plants in Tehama County, geographically and ecologically disjunct from the main range, were recorded as growing in basalt crevices.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 417.

FNA vol. 17, p. 420.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus glaucescens, M. guttatus var. glaucescens

Mimulus pardalis, M. cupriphilus, M. guttatus var. cupriphilus, M. guttatus var. pardalis

Name authority

(Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)

(Pennell) G. L. Nesom: Phytoneuron 2012-39: 44. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Erythranthe glaucescens

Erythranthe pardalis

Erythranthe glaucescens

Erythranthe pardalis