Erythranthe glaucescens |
Erythranthe michiganensis |
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shield-bract monkeyflower |
Michigan monkeyflower |
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Habit | Annuals, slender-taprooted or fibrous-rooted, rarely with runners from basal nodes. | Perennials, rhizomatous, usually producing numerous leafy stolons from basal nodes, rooting at distal nodes, sometimes forming mats. |
Stems | erect, simple or branched, terete, sometimes 4-angled distally, (5–)30–60(–80) cm, glabrous, glaucous. |
ascending-erect or basally decumbent, becoming erect in inflorescence, branched, 12–50(–70) cm, glabrous or minutely hirtellous and stipitate-glandular. |
Leaves | basal and cauline; petiole: basal and proximal cauline as long as or much longer than blade, slender, sometimes pubescent or villous, distals absent; blade palmately 3–5-veined, (proximal) ovate to ovate-elliptic or orbicular-ovate, sometimes subcordate, 10–50 mm, midcauline to distal orbicular, 5–45 mm wide, distinctly connate-perfoliate, disclike distally, base rounded to subcordate, margins: proximals denticulate to dentate or coarsely, irregularly toothed, sometimes lobed at base, distals nearly entire or toothed, teeth scattered, small, apex rounded, surfaces glabrous, glaucous. |
cauline, basal not persistent; petiole 1–5(–15) mm, mid stem and distals 0 mm; blade palmately 3–5-veined, broadly ovate to broadly ovate-elliptic or suborbicular, 8–30 mm, relatively even-sized or diminishing in size distally, bracteal reduced, slightly falcate, base truncate to cuneate, margins evenly or unevenly dentate-serrate to dentate, teeth 3–8 per side, apex usually rounded, surfaces glabrous or minutely hirtellous and stipitate-glandular. |
Flowers | herkogamous, 1–16, from distal nodes, sometimes from nearly all, chasmogamous. |
herkogamous, 2–14, mostly from distal nodes or medial to distal nodes. |
Styles | minutely hirtellous-puberulent. |
glabrous. |
Corollas | yellow, sometimes with a median splotch, abaxial limb densely dark yellow, others much lighter, throat floor and tube red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 12–23 mm, exserted 4–8 mm beyond calyx margin; limb expanded 14–36 mm. |
yellow, sometimes faintly red-spotted, bilaterally symmetric, strongly bilabiate; tube-throat cylindric-campanulate, 10–14 mm, exserted 5–8 mm beyond calyx margin; limb expanded 10–15 mm. |
Fruiting pedicels | 10–50 mm, glabrous, glaucous. |
10–25 mm, villous-glandular to minutely villosulous-glandular. |
Fruiting calyces | broadly campanulate, inflated, sagittally compressed, 7–16 mm, glabrous, glaucous, throat closing. |
cylindric-campanulate, inflated, sagittally compressed, 7–10 mm, puberulous to softly hirtellous, mixed with longer, stipitate-glandular hairs, throat not closing, adaxial lobe 2 times longer than others, slightly upcurving. |
Capsules | included, 5–11 mm. |
unknown. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28. |
= (28)30. |
Erythranthe glaucescens |
Erythranthe michiganensis |
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Phenology | Flowering Mar–May(–Jun). | Flowering Jun–Aug(–Oct). |
Habitat | Seepage areas, wet rocks, moist cliffs, pool edges, gravelly stream banks, serpentine outcrops, roadsides and roadcuts, low pastures, riparian woodlands, blue oak woodlands, chaparral, grasslands. | Cold calcareous springs, seeps, depressions, streams, alkaline shorelines at mouths of small drainages, steep moraine slopes, bluff bases, commonly in northern white cedar swamps. |
Elevation | 80–900(–1100) m. (300–3000(–3600) ft.) | 500–600 m. (1600–2000 ft.) |
Distribution |
CA
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MI |
Discussion | Plants from one locality in Butte County are unusual in producing filiform, small-leaved runners from basal cauline nodes. Erythranthe glaucescens is known only from Butte and Tehama counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Based on data from allozyme and RAPD studies, morphology, and crossing studies, A. L. Posto and L. A. Prather (2003) raised Erythranthe [Mimulus] michiganensis to specific rank. The species is endemic to a small area in the Mackinac Straits and Grand Traverse regions of Michigan; it is known to be extant at 15 sites and apparently is extinct at three previously known sites. Plants of all but one of the populations are essentially pollen-sterile and reproduce through rhizomes. However, the didynamous stamens and stigma positioned above the adaxial anther pair, along with the relatively large corollas with broadly expanded limb, are reflective of an allogamous breeding system, which seems unusual in view of its apparent sterility. The narrow geographic distribution of E. michiganensis lies within the wider range of E. geyeri, and the two are known to co-occur at two sites, apparently without hybridization or morphologically intermediate individuals. Erythranthe michiganensis (as Mimulus michiganensis) is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 417. | FNA vol. 17, p. 407. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus glaucescens, M. guttatus var. glaucescens | Mimulus glabratus subsp. michiganensis, M. glabratus var. michiganensis, M. michiganensis |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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