Erythranthe glaucescens |
Erythranthe marmorata |
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shield-bract monkeyflower |
Stanislaus monkeyflower, Whipple's monkeyflower |
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Habit | Annuals, slender-taprooted or fibrous-rooted, rarely with runners from basal nodes. | Annuals, taprooted. |
Stems | erect, simple or branched, terete, sometimes 4-angled distally, (5–)30–60(–80) cm, glabrous, glaucous. |
erect, simple or branched from base, 7–28 cm, usually moderately to densely villous-glandular, without eglandular hairs. |
Leaves | basal and cauline; petiole: basal and proximal cauline as long as or much longer than blade, slender, sometimes pubescent or villous, distals absent; blade palmately 3–5-veined, (proximal) ovate to ovate-elliptic or orbicular-ovate, sometimes subcordate, 10–50 mm, midcauline to distal orbicular, 5–45 mm wide, distinctly connate-perfoliate, disclike distally, base rounded to subcordate, margins: proximals denticulate to dentate or coarsely, irregularly toothed, sometimes lobed at base, distals nearly entire or toothed, teeth scattered, small, apex rounded, surfaces glabrous, glaucous. |
usually cauline or basal persistent; petiole: proximals to medials 7–15 mm, distals 0 mm; blade palmately (3–)5-veined, ovate or broadly ovate to elliptic-ovate or depressed-ovate, (10–)15–30 × 6–15 mm, base truncate to shallowly cuneate, margins shallowly to coarsely dentate, apex acute, surfaces usually moderately to densely villous-glandular, without eglandular hairs. |
Flowers | herkogamous, 1–16, from distal nodes, sometimes from nearly all, chasmogamous. |
herkogamous, (1–)2–6, axillary from middle to distal nodes, chasmogamous. |
Styles | minutely hirtellous-puberulent. |
glabrous or sparsely hirtellous. |
Corollas | yellow, sometimes with a median splotch, abaxial limb densely dark yellow, others much lighter, throat floor and tube red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 12–23 mm, exserted 4–8 mm beyond calyx margin; limb expanded 14–36 mm. |
yellow, throat red-spotted, abaxial limb base with a large red splotch, bilaterally symmetric, bilabiate; tube-throat narrowly cylindric-funnelform, 10–12 mm, exserted 4–5 mm beyond calyx margin; limb abruptly expanded 14–20 mm. |
Fruiting pedicels | 10–50 mm, glabrous, glaucous. |
15–45 mm, usually moderately to densely villous-glandular, without eglandular hairs. |
Fruiting calyces | broadly campanulate, inflated, sagittally compressed, 7–16 mm, glabrous, glaucous, throat closing. |
sharply nodding, usually densely purple-spotted, broadly campanulate, inflated, sagittally compressed, 9–12 mm, densely hirtellous, sometimes sparsely stipitate-glandular, densely villous at sinuses, throat closing, adaxial lobe ca. 2 times length of others. |
Capsules | included, 5–11 mm. |
included, 6–9 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28. |
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Erythranthe glaucescens |
Erythranthe marmorata |
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Phenology | Flowering Mar–May(–Jun). | Flowering Mar–May. |
Habitat | Seepage areas, wet rocks, moist cliffs, pool edges, gravelly stream banks, serpentine outcrops, roadsides and roadcuts, low pastures, riparian woodlands, blue oak woodlands, chaparral, grasslands. | Habitat unknown, not over serpentine. |
Elevation | 80–900(–1100) m. (300–3000(–3600) ft.) | 100–900 m. (300–3000 ft.) |
Distribution |
CA
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CA |
Discussion | Plants from one locality in Butte County are unusual in producing filiform, small-leaved runners from basal cauline nodes. Erythranthe glaucescens is known only from Butte and Tehama counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe marmorata is recognized by its erect, taprooted habit and annual duration, villous-glandular vestiture, ovate-petiolate leaves, flowers from middle to distal nodes, long, narrow corolla tube-throat abruptly flaring into a broad limb, abaxial middle corolla lobe with a large red splotch, and fruiting calyces dark-spotted and sharply nodding. The species is known from foothills in Calaveras, Stanislaus, and Tuolumne counties in the Stanislaus River drainage and from Amador County in the Mokelumne River drainage. A collection from Fresno County appears to be somewhat disjunct from the main range, and the plants are more densely villous than characteristic elsewhere, but their identification as E. marmorata otherwise seems secure. Erythranthe marmorata (previously identified as Mimulus whipplei) had been considered extremely rare or even perhaps extinct. See G. L. Nesom (2013d) for citations of recent collections. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 417. | FNA vol. 17, p. 416. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus glaucescens, M. guttatus var. glaucescens | Mimulus marmoratus, M. whipplei |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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