FNA: Erythranthe glaucescens vs. Erythranthe hardhamiae

	Erythranthe glaucescens	Erythranthe hardhamiae
	shield-bract monkeyflower	Hardham's monkeyflower, Santa Lucia monkeyflower
Habit	Annuals, slender-taprooted or fibrous-rooted, rarely with runners from basal nodes.	Annuals, taprooted.
Stems	erect, simple or branched, terete, sometimes 4-angled distally, (5–)30–60(–80) cm, glabrous, glaucous.	erect, simple or branched from basal nodes, (2–)3–13 cm, glabrous or minutely puberulent.
Leaves	basal and cauline; petiole: basal and proximal cauline as long as or much longer than blade, slender, sometimes pubescent or villous, distals absent; blade palmately 3–5-veined, (proximal) ovate to ovate-elliptic or orbicular-ovate, sometimes subcordate, 10–50 mm, midcauline to distal orbicular, 5–45 mm wide, distinctly connate-perfoliate, disclike distally, base rounded to subcordate, margins: proximals denticulate to dentate or coarsely, irregularly toothed, sometimes lobed at base, distals nearly entire or toothed, teeth scattered, small, apex rounded, surfaces glabrous, glaucous.	cauline, basal not persistent; petiole 0 mm; blade palmately 3-veined (in broader ones), linear to oblanceolate, 2–12 × 1–5 mm, base attenuate, margins entire, sometimes toothed, apex acute, surfaces glabrous or minutely puberulent.
Flowers	herkogamous, 1–16, from distal nodes, sometimes from nearly all, chasmogamous.	herkogamous, 1–12, from distal or medial to distal nodes.
Styles	minutely hirtellous-puberulent.	glabrous.
Corollas	yellow, sometimes with a median splotch, abaxial limb densely dark yellow, others much lighter, throat floor and tube red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 12–23 mm, exserted 4–8 mm beyond calyx margin; limb expanded 14–36 mm.	pink to purple, abaxial limb with 2 yellow palate ridges, bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 5–10 mm, exserted 2–5 mm beyond calyx margin; limb expanded 7–13 mm, lobes notched, abaxial limb sparsely bearded.
Fruiting pedicels	10–50 mm, glabrous, glaucous.	ascending to often spreading horizontally, 10–60 mm.
Fruiting calyces	broadly campanulate, inflated, sagittally compressed, 7–16 mm, glabrous, glaucous, throat closing.	becoming reddish, sometimes red-spotted, campanulate to cylindric, 4–6 mm, margins distinctly toothed or lobed, glabrous or minutely puberulent, ribs weak, lobes pronounced, erect, margins glabrous.
Capsules	included, 5–11 mm.	included, 4–5 mm.
Anthers	included, glabrous.	included or slightly exserted, glabrous.
Stigmas		equal in length to corolla tube or exserted.
2n	= 28.

	Erythranthe glaucescens	Erythranthe hardhamiae
Phenology	Flowering Mar–May(–Jun).	Flowering Mar–Apr.
Habitat	Seepage areas, wet rocks, moist cliffs, pool edges, gravelly stream banks, serpentine outcrops, roadsides and roadcuts, low pastures, riparian woodlands, blue oak woodlands, chaparral, grasslands.	Sandy soils near sandstone outcrops and chaparral.
Elevation	80–900(–1100) m. (300–3000(–3600) ft.)	300–800 m. (1000–2600 ft.)
Distribution	CA [WildflowerSearch map]	CA
Discussion	Plants from one locality in Butte County are unusual in producing filiform, small-leaved runners from basal cauline nodes. Erythranthe glaucescens is known only from Butte and Tehama counties. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Erythranthe hardhamiae is endemic to the central coast region in Monterey and San Luis Obispo counties. The species was previously included in E. palmeri but can be distinguished by having a wider limb (16–25 mm) than E. palmeri (8–15 mm) and pale pink flowers with a broad yellow palate and orifice. In contrast, E. palmeri has deep pink flowers with two yellow ridges on the palate. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 417.	FNA vol. 17, p. 388.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus glaucescens, M. guttatus var. glaucescens
Name authority	(Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)	N. S. Fraga: Aliso 30: 64, figs. 23–25. (2012)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe hardhamiae

shield-bract monkeyflower

Hardham's monkeyflower, Santa Lucia monkeyflower

Habit

Annuals, slender-taprooted or fibrous-rooted, rarely with runners from basal nodes.

Annuals, taprooted.

Stems

erect, simple or branched, terete, sometimes 4-angled distally, (5–)30–60(–80) cm, glabrous, glaucous.

erect, simple or branched from basal nodes, (2–)3–13 cm, glabrous or minutely puberulent.

Leaves

basal and cauline;

petiole: basal and proximal cauline as long as or much longer than blade, slender, sometimes pubescent or villous, distals absent;

blade palmately 3–5-veined, (proximal) ovate to ovate-elliptic or orbicular-ovate, sometimes subcordate, 10–50 mm, midcauline to distal orbicular, 5–45 mm wide, distinctly connate-perfoliate, disclike distally, base rounded to subcordate, margins: proximals denticulate to dentate or coarsely, irregularly toothed, sometimes lobed at base, distals nearly entire or toothed, teeth scattered, small, apex rounded, surfaces glabrous, glaucous.

cauline, basal not persistent;

petiole 0 mm;

blade palmately 3-veined (in broader ones), linear to oblanceolate, 2–12 × 1–5 mm, base attenuate, margins entire, sometimes toothed, apex acute, surfaces glabrous or minutely puberulent.

Flowers

herkogamous, 1–16, from distal nodes, sometimes from nearly all, chasmogamous.

herkogamous, 1–12, from distal or medial to distal nodes.

Styles

minutely hirtellous-puberulent.

glabrous.

Corollas

yellow, sometimes with a median splotch, abaxial limb densely dark yellow, others much lighter, throat floor and tube red-dotted, bilaterally symmetric, bilabiate;

tube-throat funnelform, 12–23 mm, exserted 4–8 mm beyond calyx margin;

limb expanded 14–36 mm.

pink to purple, abaxial limb with 2 yellow palate ridges, bilaterally symmetric, strongly bilabiate;

tube-throat funnelform, 5–10 mm, exserted 2–5 mm beyond calyx margin;

limb expanded 7–13 mm, lobes notched, abaxial limb sparsely bearded.

Fruiting pedicels

10–50 mm, glabrous, glaucous.

ascending to often spreading horizontally, 10–60 mm.

Fruiting calyces

broadly campanulate, inflated, sagittally compressed, 7–16 mm, glabrous, glaucous, throat closing.

becoming reddish, sometimes red-spotted, campanulate to cylindric, 4–6 mm, margins distinctly toothed or lobed, glabrous or minutely puberulent, ribs weak, lobes pronounced, erect, margins glabrous.

Capsules

included, 5–11 mm.

included, 4–5 mm.

Anthers

included, glabrous.

included or slightly exserted, glabrous.

Stigmas

equal in length to corolla tube or exserted.

= 28.

Erythranthe glaucescens

Erythranthe hardhamiae

Phenology

Flowering Mar–May(–Jun).

Flowering Mar–Apr.

Habitat

Seepage areas, wet rocks, moist cliffs, pool edges, gravelly stream banks, serpentine outcrops, roadsides and roadcuts, low pastures, riparian woodlands, blue oak woodlands, chaparral, grasslands.

Sandy soils near sandstone outcrops and chaparral.

Elevation

80–900(–1100) m. (300–3000(–3600) ft.)

300–800 m. (1000–2600 ft.)

Distribution

[WildflowerSearch map]

Discussion

Plants from one locality in Butte County are unusual in producing filiform, small-leaved runners from basal cauline nodes. Erythranthe glaucescens is known only from Butte and Tehama counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe hardhamiae is endemic to the central coast region in Monterey and San Luis Obispo counties. The species was previously included in E. palmeri but can be distinguished by having a wider limb (16–25 mm) than E. palmeri (8–15 mm) and pale pink flowers with a broad yellow palate and orifice. In contrast, E. palmeri has deep pink flowers with two yellow ridges on the palate.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 417.

FNA vol. 17, p. 388.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus glaucescens, M. guttatus var. glaucescens

Name authority

(Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)

N. S. Fraga: Aliso 30: 64, figs. 23–25. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Erythranthe glaucescens

Erythranthe hardhamiae

Erythranthe glaucescens

Erythranthe hardhamiae