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shield-bract monkeyflower

Rocky Mountain or petiole-purse monkeyflower

Habit Annuals, slender-taprooted or fibrous-rooted, rarely with runners from basal nodes. Annuals, taprooted.
Stems

erect, simple or branched, terete, sometimes 4-angled distally, (5–)30–60(–80) cm, glabrous, glaucous.

erect, straight at nodes, simple, 1–10 cm, glabrous.

Leaves

basal and cauline;

petiole: basal and proximal cauline as long as or much longer than blade, slender, sometimes pubescent or villous, distals absent;

blade palmately 3–5-veined, (proximal) ovate to ovate-elliptic or orbicular-ovate, sometimes subcordate, 10–50 mm, midcauline to distal orbicular, 5–45 mm wide, distinctly connate-perfoliate, disclike distally, base rounded to subcordate, margins: proximals denticulate to dentate or coarsely, irregularly toothed, sometimes lobed at base, distals nearly entire or toothed, teeth scattered, small, apex rounded, surfaces glabrous, glaucous.

cauline;

petiole 2–3 mm, laterally compressed, base deeply saccate, usually containing a lenticular propagule;

blade emerging from bulbils, palmately veined, elliptic-ovate to ovate, 2–8(–10) × 2–5(–7) mm, base truncate to shallowly cordate, margins entire or remotely denticulate, apex obtuse to rounded, surfaces glabrous.

Flowers

herkogamous, 1–16, from distal nodes, sometimes from nearly all, chasmogamous.

herkogamous, (1 or)2–12, from medial or medial to distal nodes.

Styles

minutely hirtellous-puberulent.

glabrous.

Corollas

yellow, sometimes with a median splotch, abaxial limb densely dark yellow, others much lighter, throat floor and tube red-dotted, bilaterally symmetric, bilabiate;

tube-throat funnelform, 12–23 mm, exserted 4–8 mm beyond calyx margin;

limb expanded 14–36 mm.

yellow, palate yellow, not spotted or striped, bilaterally symmetric, weakly bilabiate;

tube-throat broadly cylindric-funnelform, 3–4 mm, exserted beyond calyx margin;

lobes subequal, oblong-obovate, throat open, palate puberulent, abaxial ridges low.

Fruiting pedicels

10–50 mm, glabrous, glaucous.

4–6 mm, slightly longer than calyx, glabrous.

Fruiting calyces

broadly campanulate, inflated, sagittally compressed, 7–16 mm, glabrous, glaucous, throat closing.

strongly angled, subcampanulate, weakly inflated, 3–4 mm, margins distinctly toothed or lobed, glabrous, lobes pronounced, erect, incurved-triangular.

Capsules

included, 5–11 mm.

unknown.

Anthers

included, glabrous.

included, glabrous.

2n

= 28.

= 16.

Erythranthe glaucescens

Erythranthe gemmipara

Phenology Flowering Mar–May(–Jun). Flowering Jul–Aug(–Sep).
Habitat Seepage areas, wet rocks, moist cliffs, pool edges, gravelly stream banks, serpentine outcrops, roadsides and roadcuts, low pastures, riparian woodlands, blue oak woodlands, chaparral, grasslands. Granitic seeps, thin soils over bedrock cliff bases, crevices, ledges, talus, among rocks and boulders, Douglas fir, spruce-fir, and aspen forests.
Elevation 80–900(–1100) m. (300–3000(–3600) ft.) 2600–3700 m. (8500–12100 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
from FNA
CO
Discussion

Plants from one locality in Butte County are unusual in producing filiform, small-leaved runners from basal cauline nodes. Erythranthe glaucescens is known only from Butte and Tehama counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe gemmipara is known only from Grand, Jefferson, Larimer, and Park counties in north-central Colorado. Flowers in this species are uncommon, and seed set has not been observed in natural populations; reproduction in nature appears to be solely asexual via overwintering propagules (bulbils) formed in leaf axils. Two meristems are initiated in each axil. The proximal meristem produces a pair of starch-thickened storage leaves, a rudimentary axis, and a distal pair of preformed leaf primordia that enclose the shoot apical meristem. Root primordia are present within the first node of the bulbil. The petiole of the subtending leaf expands laterally and folds adaxially to enclose the developing bulbil, and entangled trichomes along the petiole margins secure it following leaf abscission and dispersal. The leaf blades commonly are deciduous, leaving the bulbil still attached (M. R. Beardsley 1997).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 417. FNA vol. 17, p. 395.
Parent taxa Phrymaceae > Erythranthe Phrymaceae > Erythranthe
Sibling taxa
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms Mimulus glaucescens, M. guttatus var. glaucescens Mimulus gemmiparus
Name authority (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) (W. A. Weber) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 37. (2012)
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