Erythranthe glaucescens |
Erythranthe floribunda |
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shield-bract monkeyflower |
many-flower monkey-flower, purple-stem monkey-flower |
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Habit | Annuals, slender-taprooted or fibrous-rooted, rarely with runners from basal nodes. | Annuals, fibrous-rooted or filiform-taprooted. |
Stems | erect, simple or branched, terete, sometimes 4-angled distally, (5–)30–60(–80) cm, glabrous, glaucous. |
erect to decumbent, sometimes procumbent-trailing, straight or geniculate at nodes, simple or many-branched, 3–22(–40) cm, villous-glandular, hairs greatly variable in length and density, gland-tipped, sometimes 0.2–0.5 mm, sparsely stipitate-glandular. |
Leaves | basal and cauline; petiole: basal and proximal cauline as long as or much longer than blade, slender, sometimes pubescent or villous, distals absent; blade palmately 3–5-veined, (proximal) ovate to ovate-elliptic or orbicular-ovate, sometimes subcordate, 10–50 mm, midcauline to distal orbicular, 5–45 mm wide, distinctly connate-perfoliate, disclike distally, base rounded to subcordate, margins: proximals denticulate to dentate or coarsely, irregularly toothed, sometimes lobed at base, distals nearly entire or toothed, teeth scattered, small, apex rounded, surfaces glabrous, glaucous. |
cauline, basal mostly deciduous by flowering; petiole 1–12 mm; blade pinnately to subpalmately veined, ovate, (3–)8–25(–35) × (1–)5–18(–26) mm, base cuneate to truncate or cordate, margins serrate to sparsely dentate, apex acute, surfaces villous-glandular, hairs greatly variable in length and density, gland-tipped, sometimes 0.2–0.5 mm, sparsely stipitate-glandular. |
Flowers | herkogamous, 1–16, from distal nodes, sometimes from nearly all, chasmogamous. |
plesiogamous, 1–20, from proximal to distal nodes. |
Styles | minutely hirtellous-puberulent. |
glabrous. |
Corollas | yellow, sometimes with a median splotch, abaxial limb densely dark yellow, others much lighter, throat floor and tube red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 12–23 mm, exserted 4–8 mm beyond calyx margin; limb expanded 14–36 mm. |
yellow, abaxial limb red-dotted, bilaterally symmetric, weakly bilabiate; tube-throat funnelform-cylindric, (4–)5–10 mm, exserted slightly beyond calyx margin or not; limb expanded 3–4 mm diam., lobes usually oblong, apex notched. |
Fruiting pedicels | 10–50 mm, glabrous, glaucous. |
5–20(–26) mm, villous-glandular, hairs greatly variable in length and density, gland-tipped, sometimes 0.2–0.5 mm, sparsely stipitate-glandular. |
Fruiting calyces | broadly campanulate, inflated, sagittally compressed, 7–16 mm, glabrous, glaucous, throat closing. |
greenish or purplish to red-dotted, cylindric, ± inflated, 4–7 mm, margins distinctly toothed or lobed, villous-glandular, lobes pronounced, erect. |
Capsules | included, 5–11 mm. |
included, 4–7 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28. |
= 32. |
Erythranthe glaucescens |
Erythranthe floribunda |
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Phenology | Flowering Mar–May(–Jun). | Flowering (May–)Jun–Aug(–Sep). |
Habitat | Seepage areas, wet rocks, moist cliffs, pool edges, gravelly stream banks, serpentine outcrops, roadsides and roadcuts, low pastures, riparian woodlands, blue oak woodlands, chaparral, grasslands. | Under overhangs, moist roofs of caves, wet rock crevices, cliff faces, wet cliff bases, below waterfalls, seeps, springs, humus and moist soils over rocks and slabs, moist slopes, ditches and pond edges, wet edges of creeks and rivers, drying mud on margins of wetland depressions, creek beds, wet or swampy meadows, along trails, in lodgepole pine, ponderosa pine, ponderosa pine-Douglas fir, and spruce-fir woodlands. |
Elevation | 80–900(–1100) m. (300–3000(–3600) ft.) | (100–)1800–2600(–3100) m. ((300–)5900–8500(–10200) ft.) |
Distribution |
CA
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AR; AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; Mexico (Baja California, Baja California Sur, Chihuahua, Sinaloa, Sonora)
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Discussion | Plants from one locality in Butte County are unusual in producing filiform, small-leaved runners from basal cauline nodes. Erythranthe glaucescens is known only from Butte and Tehama counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Some plants identified here as Erythranthe floribunda in Arizona and southwestern New Mexico are distinctive in their prominently inflated calyces, sessile to subsessile leaves with attenuate bases and palmately three- to five-veined venation, and much-elongated pedicels (20–43 mm); numerous intermediates in Arizona make it difficult to conclude that the variants represent an entity discontinuous from plants of typical morphology. The variant morphology has not been observed among Mexican populations. Further discussion of this situation was given by G. L. Nesom (2012h). Erythranthe floribunda has been documented from 12 counties in northern Arkansas (Carroll, Cleburne, Crawford, Franklin, Izard, Johnson, Logan, Newton, Pope, Searcy, Stone, and Washington), where it occurs at 300–500 m. The unpublished name Mimulus floribundus subsp. moorei Iltis appears in various checklists in reference to the Arkansas plants, but there appears to be no basis for treating them as distinct from the rest of the species. Elsewhere in the main range (western states), scattered variants extremely reduced in size, leaves, flowers, and overall stature appear to be at the lower limits of the species rather than taxonomically distinct. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 417. | FNA vol. 17, p. 404. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus glaucescens, M. guttatus var. glaucescens | Mimulus floribundus, M. deltoideus, M. floribundus var. membranaceus, M. membranaceus, M. peduncularis, M. serotinus |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) | (Lindley) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) |
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