Erythranthe glaucescens |
Erythranthe arenaria |
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shield-bract monkeyflower |
sand-loving monkeyflower |
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Habit | Annuals, slender-taprooted or fibrous-rooted, rarely with runners from basal nodes. | Annuals, fibrous-rooted or filiform-taprooted. |
Stems | erect, simple or branched, terete, sometimes 4-angled distally, (5–)30–60(–80) cm, glabrous, glaucous. |
erect to ascending, straight or geniculate at nodes, simple or branched, 5–20 cm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
Leaves | basal and cauline; petiole: basal and proximal cauline as long as or much longer than blade, slender, sometimes pubescent or villous, distals absent; blade palmately 3–5-veined, (proximal) ovate to ovate-elliptic or orbicular-ovate, sometimes subcordate, 10–50 mm, midcauline to distal orbicular, 5–45 mm wide, distinctly connate-perfoliate, disclike distally, base rounded to subcordate, margins: proximals denticulate to dentate or coarsely, irregularly toothed, sometimes lobed at base, distals nearly entire or toothed, teeth scattered, small, apex rounded, surfaces glabrous, glaucous. |
basal and cauline; petiole 0 mm or proximals 1–3(–5) mm; blade 1-veined or palmately 3-veined, elliptic to narrowly elliptic, ovate-elliptic, or ovate-lanceolate, 5–12(–17) × 3–7 mm, base rounded to cuneate-attenuate, margins entire or sparsely dentate to serrate, apex acuminate to acute or obtuse, surfaces villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
Flowers | herkogamous, 1–16, from distal nodes, sometimes from nearly all, chasmogamous. |
herkogamous, 1–22, from proximal to distal nodes. |
Styles | minutely hirtellous-puberulent. |
glabrous. |
Corollas | yellow, sometimes with a median splotch, abaxial limb densely dark yellow, others much lighter, throat floor and tube red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 12–23 mm, exserted 4–8 mm beyond calyx margin; limb expanded 14–36 mm. |
yellow, abaxial limb red-dotted, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 9–12(–14) mm, exserted beyond calyx margin; lobes broadly obovate, apex rounded. |
Fruiting pedicels | 10–50 mm, glabrous, glaucous. |
divergent-arcuate, 10–23 mm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
Fruiting calyces | broadly campanulate, inflated, sagittally compressed, 7–16 mm, glabrous, glaucous, throat closing. |
usually red-dotted, narrowly campanulate, not or weakly inflated, 5–7(–9) mm, margins distinctly toothed or lobed, villous-glandular, ribs angled, lobes pronounced, erect. |
Capsules | included, 5–11 mm. |
included, 4–7 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 28. |
= 32. |
Erythranthe glaucescens |
Erythranthe arenaria |
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Phenology | Flowering Mar–May(–Jun). | Flowering May–Sep. |
Habitat | Seepage areas, wet rocks, moist cliffs, pool edges, gravelly stream banks, serpentine outcrops, roadsides and roadcuts, low pastures, riparian woodlands, blue oak woodlands, chaparral, grasslands. | Sandy flats, bars, gullies, washes, trails, roadcuts, seasonal creek beds and drainages, rocky slopes, seepy loam, ditches, lake edges, meadows, openings in pine-fir and pine-oak woodlands. |
Elevation | 80–900(–1100) m. (300–3000(–3600) ft.) | (100–)500–2600(–2800) m. ((300–)1600–8500(–9200) ft.) |
Distribution |
CA
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CA
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Discussion | Plants from one locality in Butte County are unusual in producing filiform, small-leaved runners from basal cauline nodes. Erythranthe glaucescens is known only from Butte and Tehama counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe arenaria is known from a cluster of six counties of the central Sierra Nevada: Fresno, Madera, Mariposa, Merced, Tulare, and Tuolumne. Most plants of Erythranthe arenaria have relatively even-sized cauline leaves, all sessile to proximally subsessile. Plants in the Yosemite area with persistent basal leaves that are short-petiolate, ovate with a cuneate base, and relatively larger than the more distal cauline ones, and possibly related to E. arenaria, have been named M. floribundus var. subulatus. These might be construed as showing the influence of E. geniculata, but that species occurs only at the lower range of elevation of E. arenaria, while plants referable to Mimulus floribundus var. subulatus occur at least to 2300 m and also have the erect habit characteristic of E. arenaria. These variants should be investigated, especially in the Yosemite area where they appear to be relatively common, with the possibility that they indeed represent a distinct entity. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 417. | FNA vol. 17, p. 403. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus glaucescens, M. guttatus var. glaucescens | Mimulus arenarius, M. floribundus var. subulatus, M. multiflorus, M. subulatus, M. trisulcatus |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) | (A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) |
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