Erythranthe geyeri |
Erythranthe utahensis |
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Geyer's monkeyflower, mimule de James |
Utah monkeyflower |
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Habit | Perennials, rhizomatous, rooting at nodes. | Perennials, rhizomatous, rooting at nodes, rhizomes filiform. |
Stems | decumbent-ascending to ascending or erect-ascending, branched, (3–)10–40 cm, glabrous. |
erect, sometimes decumbent-ascending proximally, simple or few-branched, 20–50 cm, glabrous or sparsely stipitate-glandular in inflorescence. |
Leaves | cauline, basal not persistent; petiole 3–10(–20) mm or 0 mm distally; blade palmately 3–5-veined, suborbicular to depressed-ovate or broadly elliptic-ovate to reniform, 6–25 mm, relatively even-sized or largest often at mid stem, bracteal reduced, base cuneate to truncate or subcordate, margins shallowly dentate to crenate-dentate, teeth 3–7(–10) per side, apex rounded, adaxial surface of distals sparsely short villous-glandular or glabrous. |
basal and cauline, even-sized or largest near mid stem; petiole 0 mm or proximalmost 2–10 mm; blade palmately 3–5-veined, orbicular or suborbicular to broadly elliptic, broadly ovate, or depressed-ovate, 20–40(–75) × 12–35(–40) mm, base usually truncate to broadly cuneate, margins entire or subentire to mucronulate, shallowly dentate, or denticulate, apex rounded, surfaces glabrous or glabrate to sparsely stipitate-glandular and sparsely pilose, hairs thin-walled, abaxial often glaucous. |
Flowers | plesiogamous, 2–8(–12), from distal nodes, sometimes from most nodes, very loosely racemose. |
herkogamous, 6–16, from proximal to distal nodes, in a loose raceme, distal bracts becoming much reduced. |
Styles | glabrous. |
hirtellous. |
Corollas | yellow, sparsely red-dotted or not, bilaterally symmetric, weakly bilabiate; tube-throat cylindric-funnelform, 6–8 mm, exserted 1–3 mm beyond calyx margin; limb expanded 5–8 mm. |
yellow, abaxial limb prominently darker yellow, sparsely purple-spotted, bilaterally symmetric, weakly bilabiate; tube-throat narrowly funnelform to broadly cylindric, 10–15 mm, exserted 5–8 mm beyond calyx margin; limb expanded 12–20 mm. |
Fruiting pedicels | 18–30 mm, sparsely short villous-glandular or glabrous. |
(25–)40–75 mm, stipitate-glandular to short villous, hairs gland-tipped. |
Fruiting calyces | obtriangular to broadly obtriangular or deeply cupulate, inflated, sagittally compressed, (7–)8–12 mm, sparsely short villous-glandular or glabrous, throat not closing, lateral lobes shallowly convex-mucronulate, adaxial ovate with apex rounded. |
broadly ovate-cylindric, inflated, sagittally compressed, (10–)11–17(–20) mm, stipitate-glandular or minutely hirtellous or a mixture, hairs sometimes also longer, thin-walled, eglandular or glandular, throat not closing, adaxial lobe slightly longer, triangular-blunt. |
Capsules | included, (4.5–)5–8 mm. |
included, 5–7 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 30. |
= 28, 30. |
Erythranthe geyeri |
Erythranthe utahensis |
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Phenology | Flowering May–Aug(–Oct). | Flowering (Feb–)May–Aug(–Oct). |
Habitat | Edges of flowing streams, marsh edges, drainage ditches, seepage areas, springs, muddy or moist banks. | Drainage ditches, springs, seeps, wet meadows, margins of ponds and small streams, marshy areas. |
Elevation | 200–2500 m. (700–8200 ft.) | 1400–2500 m. (4600–8200 ft.) |
Distribution |
AZ; CO; IA; IL; KS; MI; MN; MO; NE; NM; OK; PA; SD; TX; WI; WY; AB; MB; ON; QC; SK; Mexico (Chihuahua, Coahuila, Distrito Federal, Durango, Hidalgo, Nuevo León, México, Querétaro, San Luis Potosí, Sonora, Veracruz, Zacatecas)
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CA; CO; NV; UT |
Discussion | Erythranthe geyeri has commonly been regarded as conspecific with E. glabrata (Kunth) G. L. Nesom (as Mimulus glabratus var. jamesii), but typical E. glabrata has a different chromosome number and distinct morphology and its range does not reach the United States. In Mexico, the two species are broadly sympatric without intermediates. An allozyme study of the M. glabratus complex (R. K. Vickery 1990) indicated that the Great Plains populations of E. geyeri are distinct from those in New Mexico and Mexico, corresponding to a difference in pedicel vestiture. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe utahensis is characterized by its erect stems, prolifically produced filiform rhizomes, basal leaves short-petiolate to subsessile and cauline sessile, blades suborbicular to broadly ovate or depressed ovate with thin-walled villous-glandular hairs on both surfaces, margins subentire, proximal pedicels elongating to 75 mm, and calyces open at maturity. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 406. | FNA vol. 17, p. 410. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus geyeri, M. glabratus var. fremontii, M. glabratus var. jamesii, M. glabratus var. oklahomensis, M. jamesii, M. jamesii var. fremontii | Mimulus glabratus subsp. utahensis |
Name authority | (Torrey) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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