Erythranthe geyeri |
Erythranthe percaulis |
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Geyer's monkeyflower, mimule de James |
serpentine canyon monkeyflower |
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Habit | Perennials, rhizomatous, rooting at nodes. | Annuals, fibrous-rooted. |
Stems | decumbent-ascending to ascending or erect-ascending, branched, (3–)10–40 cm, glabrous. |
erect or slightly ascending from base, often purplish, simple or few-branched from basal nodes, 7–28 cm, glabrous. |
Leaves | cauline, basal not persistent; petiole 3–10(–20) mm or 0 mm distally; blade palmately 3–5-veined, suborbicular to depressed-ovate or broadly elliptic-ovate to reniform, 6–25 mm, relatively even-sized or largest often at mid stem, bracteal reduced, base cuneate to truncate or subcordate, margins shallowly dentate to crenate-dentate, teeth 3–7(–10) per side, apex rounded, adaxial surface of distals sparsely short villous-glandular or glabrous. |
basal and cauline; petiole: basal and proximalmost cauline 5–10 mm or cauline 0 mm; blade often spreading at right angles to stem, purple, palmately 3–5-veined, narrowly ovate, rhombic-elliptic, ovate to lanceolate, or elliptic-lanceolate to ovate or oblong-ovate, 7–10 mm, cauline even-sized or slightly smaller distally, 4–10 mm, base truncate to attenuate, margins entire or proximals shallowly sinuate, serrations 2–4, shallow, apex rounded, surfaces glabrous. |
Flowers | plesiogamous, 2–8(–12), from distal nodes, sometimes from most nodes, very loosely racemose. |
herkogamous, 8–12, usually on distal 2/3 of stem, not clustered, chasmogamous. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, sparsely red-dotted or not, bilaterally symmetric, weakly bilabiate; tube-throat cylindric-funnelform, 6–8 mm, exserted 1–3 mm beyond calyx margin; limb expanded 5–8 mm. |
yellow, throat floor with a few red dots, proximal middle lip base with a larger red splotch, bilaterally symmetric, bilabiate; tube-throat cylindric-funnelform, 4–6 mm, exserted 2–3 mm beyond calyx margin; limb expanded 4–5 mm, palate ridges yellow, densely hairy. |
Fruiting pedicels | 18–30 mm, sparsely short villous-glandular or glabrous. |
15–35 mm, glabrous. |
Fruiting calyces | obtriangular to broadly obtriangular or deeply cupulate, inflated, sagittally compressed, (7–)8–12 mm, sparsely short villous-glandular or glabrous, throat not closing, lateral lobes shallowly convex-mucronulate, adaxial ovate with apex rounded. |
sharply wing-angled, urceolate to urceolate-campanulate, inflated, sagittally compressed, (4–)5–6 mm, glabrous, throat closing. |
Capsules | included, (4.5–)5–8 mm. |
included, 3 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 30. |
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Erythranthe geyeri |
Erythranthe percaulis |
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Phenology | Flowering May–Aug(–Oct). | Flowering May–Jun. |
Habitat | Edges of flowing streams, marsh edges, drainage ditches, seepage areas, springs, muddy or moist banks. | Soil pockets, crevices, and boulders on serpentine cliffs, slopes, and roadcuts. |
Elevation | 200–2500 m. (700–8200 ft.) | 2800 m. (9200 ft.) |
Distribution |
AZ; CO; IA; IL; KS; MI; MN; MO; NE; NM; OK; PA; SD; TX; WI; WY; AB; MB; ON; QC; SK; Mexico (Chihuahua, Coahuila, Distrito Federal, Durango, Hidalgo, Nuevo León, México, Querétaro, San Luis Potosí, Sonora, Veracruz, Zacatecas)
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CA |
Discussion | Erythranthe geyeri has commonly been regarded as conspecific with E. glabrata (Kunth) G. L. Nesom (as Mimulus glabratus var. jamesii), but typical E. glabrata has a different chromosome number and distinct morphology and its range does not reach the United States. In Mexico, the two species are broadly sympatric without intermediates. An allozyme study of the M. glabratus complex (R. K. Vickery 1990) indicated that the Great Plains populations of E. geyeri are distinct from those in New Mexico and Mexico, corresponding to a difference in pedicel vestiture. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe percaulis was described from only the type collection from Serpentine Canyon of the Feather River in Plumas County, but the type locality has recently been relocated and the population determined to comprise many thousands of individuals (S. Schoenig 2016). Plants are characterized by their completely glabrous vestiture, terete and mostly simple stems, small leaves on relatively widely spaced nodes, small calyces, and small, yellow corollas with herkogamous arrangement of stigma and anthers. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 406. | FNA vol. 17, p. 418. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus geyeri, M. glabratus var. fremontii, M. glabratus var. jamesii, M. glabratus var. oklahomensis, M. jamesii, M. jamesii var. fremontii | |
Name authority | (Torrey) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) | G. L. Nesom: Phytoneuron 2013-70: 1, figs. 1–5. (2013) |
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