Erythranthe geyeri |
Erythranthe inflatula |
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Geyer's monkeyflower, mimule de James |
disappearing monkey-flower, ephemeral monkeyflower |
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Habit | Perennials, rhizomatous, rooting at nodes. | Annuals, fibrous-rooted or filiform-taprooted. |
Stems | decumbent-ascending to ascending or erect-ascending, branched, (3–)10–40 cm, glabrous. |
erect to ascending, straight or geniculate at nodes, simple or branched at proximal and medial nodes, 6–20(–25) cm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
Leaves | cauline, basal not persistent; petiole 3–10(–20) mm or 0 mm distally; blade palmately 3–5-veined, suborbicular to depressed-ovate or broadly elliptic-ovate to reniform, 6–25 mm, relatively even-sized or largest often at mid stem, bracteal reduced, base cuneate to truncate or subcordate, margins shallowly dentate to crenate-dentate, teeth 3–7(–10) per side, apex rounded, adaxial surface of distals sparsely short villous-glandular or glabrous. |
usually cauline, basal usually deciduous by flowering; petiole: proximals 1–3 mm, distals 0 mm; blade palmately 3–5-veined, narrowly ovate or narrowly lanceolate to elliptic or elliptic-lanceolate, largest 8–18(–30) × (1–)3–7 mm, relatively even-sized, or slightly reduced distally, base attenuate to obtuse or rounded, margins entire, mucronulate, or denticulate, apex acute to obtuse, surfaces minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
Flowers | plesiogamous, 2–8(–12), from distal nodes, sometimes from most nodes, very loosely racemose. |
plesiogamous, 10–20, from medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, sparsely red-dotted or not, bilaterally symmetric, weakly bilabiate; tube-throat cylindric-funnelform, 6–8 mm, exserted 1–3 mm beyond calyx margin; limb expanded 5–8 mm. |
yellow to pale yellow, sparsely red-spotted or not, bilaterally symmetric, weakly bilabiate; tube-throat cylindric, 5–8 mm, exserted 1–3 mm beyond calyx margin; limb barely widened, lobes broadly obovate, apex rounded or mucronate. |
Fruiting pedicels | 18–30 mm, sparsely short villous-glandular or glabrous. |
straight, 7–18 mm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
Fruiting calyces | obtriangular to broadly obtriangular or deeply cupulate, inflated, sagittally compressed, (7–)8–12 mm, sparsely short villous-glandular or glabrous, throat not closing, lateral lobes shallowly convex-mucronulate, adaxial ovate with apex rounded. |
winged, plicate-angled, maturing ovoid-ellipsoid to campanulate or broadly urceolate, distinctly inflated, 7–11 mm, margins distinctly toothed or lobed, sparsely, minutely hirtellous, eglandular, lobes pronounced, erect. |
Capsules | included, (4.5–)5–8 mm. |
included, 5–9 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 30. |
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Erythranthe geyeri |
Erythranthe inflatula |
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Phenology | Flowering May–Aug(–Oct). | Flowering Jun–Jul. |
Habitat | Edges of flowing streams, marsh edges, drainage ditches, seepage areas, springs, muddy or moist banks. | Drying edges, banks, and beds of summer-dry watercourses, near drying edges of small lakes or impoundments, often among rocks and shoreline detritus, occasionally in moist protected areas beneath low shrubs. |
Elevation | 200–2500 m. (700–8200 ft.) | 1200–1700 m. (3900–5600 ft.) |
Distribution |
AZ; CO; IA; IL; KS; MI; MN; MO; NE; NM; OK; PA; SD; TX; WI; WY; AB; MB; ON; QC; SK; Mexico (Chihuahua, Coahuila, Distrito Federal, Durango, Hidalgo, Nuevo León, México, Querétaro, San Luis Potosí, Sonora, Veracruz, Zacatecas)
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CA; ID; NV; OR |
Discussion | Erythranthe geyeri has commonly been regarded as conspecific with E. glabrata (Kunth) G. L. Nesom (as Mimulus glabratus var. jamesii), but typical E. glabrata has a different chromosome number and distinct morphology and its range does not reach the United States. In Mexico, the two species are broadly sympatric without intermediates. An allozyme study of the M. glabratus complex (R. K. Vickery 1990) indicated that the Great Plains populations of E. geyeri are distinct from those in New Mexico and Mexico, corresponding to a difference in pedicel vestiture. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
No natural occurrences of Erythranthe inflatula are known from Washington; the type collection from Klickitat County is from a cultivated plant. Morphological and molecular data (R. J. Meinke 1995; P. M. Beardsley et al. 2004) indicate that Erythranthe inflatula originated as a hybrid between E. breviflora and E. latidens. Its geography and biology suggest that it is reproductively stable. The putative parents are geographically and ecologically separated for most of their ranges, and the range of E. inflatula is considerably broader than the relatively small region where the parents are sympatric. In the region of sympatry, however, E. inflatula may be difficult to distinguish from one or both of its putative parents. G. L. Nesom (2012g) was not able to find morphology that would distinguish the recently described Mimulus evanescens from E. inflatula. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 406. | FNA vol. 17, p. 400. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus geyeri, M. glabratus var. fremontii, M. glabratus var. jamesii, M. glabratus var. oklahomensis, M. jamesii, M. jamesii var. fremontii | Mimulus inflatulus, M. evanescens |
Name authority | (Torrey) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) | (Suksdorf) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) |
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