Erythranthe geyeri |
Erythranthe grayi |
|
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Geyer's monkeyflower, mimule de James |
Gray's monkeyflower, mariposa monkeyflower |
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Habit | Perennials, rhizomatous, rooting at nodes. | Annuals, fibrous-rooted. |
Stems | decumbent-ascending to ascending or erect-ascending, branched, (3–)10–40 cm, glabrous. |
erect, simple or branched from base, weakly 4-angled, 8–20 cm, glabrous. |
Leaves | cauline, basal not persistent; petiole 3–10(–20) mm or 0 mm distally; blade palmately 3–5-veined, suborbicular to depressed-ovate or broadly elliptic-ovate to reniform, 6–25 mm, relatively even-sized or largest often at mid stem, bracteal reduced, base cuneate to truncate or subcordate, margins shallowly dentate to crenate-dentate, teeth 3–7(–10) per side, apex rounded, adaxial surface of distals sparsely short villous-glandular or glabrous. |
basal and cauline; petiole 0 mm; blade palmately 3–5-veined, broadly ovate, 7–18 × 5–12 mm, base rounded, margins denticulate, apex acute, surfaces glabrous, rarely sparsely puberulent. |
Flowers | plesiogamous, 2–8(–12), from distal nodes, sometimes from most nodes, very loosely racemose. |
herkogamous, 2–20, from proximal to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, sparsely red-dotted or not, bilaterally symmetric, weakly bilabiate; tube-throat cylindric-funnelform, 6–8 mm, exserted 1–3 mm beyond calyx margin; limb expanded 5–8 mm. |
rose red, throat pink lined with rose red and a yellow patch, abaxial ridges yellow, bilaterally symmetric, weakly bilabiate; tube-throat cylindric-funnelform, 8–11 mm, exserted beyond calyx margin; limb expanded 7–10 mm. |
Fruiting pedicels | 18–30 mm, sparsely short villous-glandular or glabrous. |
6–7 mm, shorter than subtending leaves, glabrous. |
Fruiting calyces | obtriangular to broadly obtriangular or deeply cupulate, inflated, sagittally compressed, (7–)8–12 mm, sparsely short villous-glandular or glabrous, throat not closing, lateral lobes shallowly convex-mucronulate, adaxial ovate with apex rounded. |
campanulate, 9–11 × 5–6 mm, margins subtruncate, glabrous, sometimes densely papillate at flowering with tiny, 1-celled, eglandular hairs, these apparently deciduous by fruiting, lobes reduced, subequal. |
Capsules | included, (4.5–)5–8 mm. |
included, 5–9 mm. |
Anthers | included, glabrous. |
included, minutely villous-hirsute. |
2n | = 30. |
|
Erythranthe geyeri |
Erythranthe grayi |
|
Phenology | Flowering May–Aug(–Oct). | Flowering May–Jul(–Oct). |
Habitat | Edges of flowing streams, marsh edges, drainage ditches, seepage areas, springs, muddy or moist banks. | Drying pond beds, creek banks, yellow pine, yellow pine-Libocedrus woodlands. |
Elevation | 200–2500 m. (700–8200 ft.) | 1000–1900 m. (3300–6200 ft.) |
Distribution |
AZ; CO; IA; IL; KS; MI; MN; MO; NE; NM; OK; PA; SD; TX; WI; WY; AB; MB; ON; QC; SK; Mexico (Chihuahua, Coahuila, Distrito Federal, Durango, Hidalgo, Nuevo León, México, Querétaro, San Luis Potosí, Sonora, Veracruz, Zacatecas)
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CA
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Discussion | Erythranthe geyeri has commonly been regarded as conspecific with E. glabrata (Kunth) G. L. Nesom (as Mimulus glabratus var. jamesii), but typical E. glabrata has a different chromosome number and distinct morphology and its range does not reach the United States. In Mexico, the two species are broadly sympatric without intermediates. An allozyme study of the M. glabratus complex (R. K. Vickery 1990) indicated that the Great Plains populations of E. geyeri are distinct from those in New Mexico and Mexico, corresponding to a difference in pedicel vestiture. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In addition to the features noted in the key and descriptions, the fruiting calyces of Erythranthe grayi are distinctly more inflated than those of E. acutidens and E. inconspicua. G. L. Nesom (2012g) maintained E. grayi as distinct from E. acutidens, relying primarily on fruiting pedicel length (see key above), but the two have nearly identical ranges (Tuolumne County south to Kern County), and study of additional collections suggests that only a single species may be represented. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 406. | FNA vol. 17, p. 382. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus geyeri, M. glabratus var. fremontii, M. glabratus var. jamesii, M. glabratus var. oklahomensis, M. jamesii, M. jamesii var. fremontii | Mimulus grayi |
Name authority | (Torrey) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) | (A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 34. (2012) |
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