Erythranthe geyeri |
Erythranthe corallina |
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Geyer's monkeyflower, mimule de James |
coralline monkeyflower |
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Habit | Perennials, rhizomatous, rooting at nodes. | Perennials, rhizomatous, rhizomes often forming a mass, usually branching, filiform. |
Stems | decumbent-ascending to ascending or erect-ascending, branched, (3–)10–40 cm, glabrous. |
usually erect to ascending-erect, few-branched, 6–25(–38) cm, moderately hirsute to hirtellous, hairs deflexed. |
Leaves | cauline, basal not persistent; petiole 3–10(–20) mm or 0 mm distally; blade palmately 3–5-veined, suborbicular to depressed-ovate or broadly elliptic-ovate to reniform, 6–25 mm, relatively even-sized or largest often at mid stem, bracteal reduced, base cuneate to truncate or subcordate, margins shallowly dentate to crenate-dentate, teeth 3–7(–10) per side, apex rounded, adaxial surface of distals sparsely short villous-glandular or glabrous. |
basal and cauline, becoming larger distally or even-sized; petiole 0 mm or proximals 1–15 mm; blade palmately 5-veined, ovate to broadly ovate, 15–45 mm, base mostly truncate to shallowly cordate, margins sharply dentate-serrate, apex obtuse, surfaces hirtellous to softly hirsute, hairs ascending, straight, dull gray, sharp-pointed, thick-walled, eglandular. |
Flowers | plesiogamous, 2–8(–12), from distal nodes, sometimes from most nodes, very loosely racemose. |
herkogamous, 1–3(–6), commonly solitary or from distal nodes. |
Styles | glabrous. |
sparsely hirtellous. |
Corollas | yellow, sparsely red-dotted or not, bilaterally symmetric, weakly bilabiate; tube-throat cylindric-funnelform, 6–8 mm, exserted 1–3 mm beyond calyx margin; limb expanded 5–8 mm. |
yellow, red-spotted, bilaterally symmetric, bilabiate; tube-throat narrowly funnelform to broadly cylindric, 13–20 mm, exserted beyond calyx margin; limb expanded 12–22 mm. |
Fruiting pedicels | 18–30 mm, sparsely short villous-glandular or glabrous. |
(10–)25–75 mm, glabrous or puberulent proximally, hairs stipitate-glandular. |
Fruiting calyces | obtriangular to broadly obtriangular or deeply cupulate, inflated, sagittally compressed, (7–)8–12 mm, sparsely short villous-glandular or glabrous, throat not closing, lateral lobes shallowly convex-mucronulate, adaxial ovate with apex rounded. |
sometimes purple-spotted, broadly cylindric-campanulate, inflated, sagittally compressed, 11–15 mm, glabrous, throat not closing, proximal lobe pair slightly upcurving. |
Capsules | included, (4.5–)5–8 mm. |
included, stipitate, 7–10 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 30. |
= 48, 56. |
Erythranthe geyeri |
Erythranthe corallina |
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Phenology | Flowering May–Aug(–Oct). | Flowering (May–)Jun–Aug. |
Habitat | Edges of flowing streams, marsh edges, drainage ditches, seepage areas, springs, muddy or moist banks. | Creek banks, moraine water courses, bogs, marshes, wet meadows, roadside ditches. |
Elevation | 200–2500 m. (700–8200 ft.) | (1400–)1700–2700(–3000) m. ((4600–)5600–8900(–9800) ft.) |
Distribution |
AZ; CO; IA; IL; KS; MI; MN; MO; NE; NM; OK; PA; SD; TX; WI; WY; AB; MB; ON; QC; SK; Mexico (Chihuahua, Coahuila, Distrito Federal, Durango, Hidalgo, Nuevo León, México, Querétaro, San Luis Potosí, Sonora, Veracruz, Zacatecas)
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CA; NV |
Discussion | Erythranthe geyeri has commonly been regarded as conspecific with E. glabrata (Kunth) G. L. Nesom (as Mimulus glabratus var. jamesii), but typical E. glabrata has a different chromosome number and distinct morphology and its range does not reach the United States. In Mexico, the two species are broadly sympatric without intermediates. An allozyme study of the M. glabratus complex (R. K. Vickery 1990) indicated that the Great Plains populations of E. geyeri are distinct from those in New Mexico and Mexico, corresponding to a difference in pedicel vestiture. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe corallina is a morphologically consistent entity that occurs only in the Sierra Nevada of California and adjacent Nevada (Washoe County and Carson City). Its chromosome number is reported as 2n = 48 and 56, compared to 2n = 28 and 56 in E. tilingii; identities of the E. corallina vouchers should be rechecked and additional counts made, since the occurrence of such wide dysploidy seems unlikely. Compared to the leaf blades of E. tilingii in the strict sense, those of E. corallina are relatively broader (broadly ovate to orbicular-ovate), the plants generally taller, and long-pedicellate flowers occasionally are produced from mid stem or even proximal nodes. The hirsutulous to hirsute vestiture of eglandular hairs on both leaf surfaces is a reliably diagnostic feature and usually easily observed with a 10× lens. Some plants of Erythranthe corallina from San Bernardino County, California, produce decumbent-ascending stems (4–10 cm) and ovate-triangular leaves (blade 5–10 × 3–6 mm), but the dense system of filiform rhizomes, flowers one to three, and hirtellous foliar vestiture serve to identify them. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 406. | FNA vol. 17, p. 410. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus geyeri, M. glabratus var. fremontii, M. glabratus var. jamesii, M. glabratus var. oklahomensis, M. jamesii, M. jamesii var. fremontii | Mimulus corallinus, M. minusculus, M. tilingii var. corallinus |
Name authority | (Torrey) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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