Erythranthe geyeri |
Erythranthe charlestonensis |
|
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Geyer's monkeyflower, mimule de James |
Charleston Mountains monkeyflower |
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Habit | Perennials, rhizomatous, rooting at nodes. | Annuals, fibrous-rooted, usually dark purplish. |
Stems | decumbent-ascending to ascending or erect-ascending, branched, (3–)10–40 cm, glabrous. |
erect, simple, weakly 4-angled, 4–16(–24) cm, slender, glabrous or sparsely villous-glandular near nodes. |
Leaves | cauline, basal not persistent; petiole 3–10(–20) mm or 0 mm distally; blade palmately 3–5-veined, suborbicular to depressed-ovate or broadly elliptic-ovate to reniform, 6–25 mm, relatively even-sized or largest often at mid stem, bracteal reduced, base cuneate to truncate or subcordate, margins shallowly dentate to crenate-dentate, teeth 3–7(–10) per side, apex rounded, adaxial surface of distals sparsely short villous-glandular or glabrous. |
basal and cauline, basal usually persistent, largest at mid stem or above, cauline relatively few on long internodes; petiole: proximals to distals 1–4 mm; blade palmately 3-veined, ovate to ovate-lanceolate, 5–16(–20) × 3–11 mm, base truncate to subcordate, margins shallowly, evenly crenulate to serrate-dentate or denticulate, apex acute to obtuse, surfaces: proximals glabrous or sparsely villous, distals and bracteals sparsely hirtellous or glabrous abaxially or along distal margin, sparsely villous adaxially, hairs vitreous, flattened, eglandular. |
Flowers | plesiogamous, 2–8(–12), from distal nodes, sometimes from most nodes, very loosely racemose. |
plesiogamous, (1–)4–7, usually from all nodes, usually beginning about mid stem, cleistogamous. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, sparsely red-dotted or not, bilaterally symmetric, weakly bilabiate; tube-throat cylindric-funnelform, 6–8 mm, exserted 1–3 mm beyond calyx margin; limb expanded 5–8 mm. |
yellow, sparsely red-dotted, bilaterally or nearly radially symmetric, bilabiate or nearly regular; tube-throat narrowly cylindric, 4–6 mm, exserted 0.5–1 mm beyond calyx margin; limb barely expanded. |
Fruiting pedicels | 18–30 mm, sparsely short villous-glandular or glabrous. |
deflexed 45–90º at calyx, 6–19 mm, longer than subtending leaves, glabrous or sparsely villous-glandular. |
Fruiting calyces | obtriangular to broadly obtriangular or deeply cupulate, inflated, sagittally compressed, (7–)8–12 mm, sparsely short villous-glandular or glabrous, throat not closing, lateral lobes shallowly convex-mucronulate, adaxial ovate with apex rounded. |
sometimes purple-dotted, broadly elliptic-ovoid, inflated, sagittally compressed, 10–13 mm, minutely hirtellous, sometimes also sparsely glandular, throat closing, adaxial lobe not distinctly longer than abaxial, not falcate. |
Capsules | included, (4.5–)5–8 mm. |
included, stipitate, 6–8 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 30. |
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Erythranthe geyeri |
Erythranthe charlestonensis |
|
Phenology | Flowering May–Aug(–Oct). | Flowering Apr–Jul(–Aug). |
Habitat | Edges of flowing streams, marsh edges, drainage ditches, seepage areas, springs, muddy or moist banks. | Grassy slopes, damp soils, moist rocks. |
Elevation | 200–2500 m. (700–8200 ft.) | (900–)1700–2400(–2800) m. ((3000–)5600–7900(–9200) ft.) |
Distribution |
AZ; CO; IA; IL; KS; MI; MN; MO; NE; NM; OK; PA; SD; TX; WI; WY; AB; MB; ON; QC; SK; Mexico (Chihuahua, Coahuila, Distrito Federal, Durango, Hidalgo, Nuevo León, México, Querétaro, San Luis Potosí, Sonora, Veracruz, Zacatecas)
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AZ; NV |
Discussion | Erythranthe geyeri has commonly been regarded as conspecific with E. glabrata (Kunth) G. L. Nesom (as Mimulus glabratus var. jamesii), but typical E. glabrata has a different chromosome number and distinct morphology and its range does not reach the United States. In Mexico, the two species are broadly sympatric without intermediates. An allozyme study of the M. glabratus complex (R. K. Vickery 1990) indicated that the Great Plains populations of E. geyeri are distinct from those in New Mexico and Mexico, corresponding to a difference in pedicel vestiture. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe charlestonensis is characterized by its annual duration, autogamous reproduction, small stature, commonly purplish color, regularly ovate, short-petiolate leaves with shallowly crenate margins and (distal leaves) sparsely villous or glabrous adaxial surfaces, hirtellous calyces without a prominently longer adaxial lobe, and lack of glandular hairs. It is endemic to the Charleston (Spring) Mountains of Clark County, Nevada, and one locality (Union Pass) in Mohave County, Arizona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 406. | FNA vol. 17, p. 423. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus geyeri, M. glabratus var. fremontii, M. glabratus var. jamesii, M. glabratus var. oklahomensis, M. jamesii, M. jamesii var. fremontii | |
Name authority | (Torrey) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) | G. L. Nesom: Phytoneuron 2012-40: 80. (2012) |
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