Erythranthe geyeri |
Erythranthe arenicola |
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Geyer's monkeyflower, mimule de James |
beach monkeyflower |
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Habit | Perennials, rhizomatous, rooting at nodes. | Annuals, fibrous-rooted or slender-taprooted, rarely rooting at nodes. |
Stems | decumbent-ascending to ascending or erect-ascending, branched, (3–)10–40 cm, glabrous. |
erect, rarely prostrate to prostrate-ascending, few-branched, 3–17 cm, moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular. |
Leaves | cauline, basal not persistent; petiole 3–10(–20) mm or 0 mm distally; blade palmately 3–5-veined, suborbicular to depressed-ovate or broadly elliptic-ovate to reniform, 6–25 mm, relatively even-sized or largest often at mid stem, bracteal reduced, base cuneate to truncate or subcordate, margins shallowly dentate to crenate-dentate, teeth 3–7(–10) per side, apex rounded, adaxial surface of distals sparsely short villous-glandular or glabrous. |
basal and cauline; petiole: basal 2–8 mm or mid and distals absent; blade palmately 3–5-veined, suborbicular to broadly ovate or depressed-ovate, 5–17 × 6–15 mm, base truncate or truncate-cuneate to subcordate, margins subentire or crenulate, apex rounded to obtuse, surfaces moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular. |
Flowers | plesiogamous, 2–8(–12), from distal nodes, sometimes from most nodes, very loosely racemose. |
herkogamous, 1–6, at distal nodes, chasmogamous. |
Styles | glabrous. |
hirtellous. |
Corollas | yellow, sparsely red-dotted or not, bilaterally symmetric, weakly bilabiate; tube-throat cylindric-funnelform, 6–8 mm, exserted 1–3 mm beyond calyx margin; limb expanded 5–8 mm. |
yellow, red-dotted, bilaterally symmetric, bilabiate; tube-throat funnelform, 11–20 mm, exserted 4–8 mm beyond calyx margin; limb expanded 10–18 mm. |
Fruiting pedicels | 18–30 mm, sparsely short villous-glandular or glabrous. |
9–17 mm, moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular. |
Fruiting calyces | obtriangular to broadly obtriangular or deeply cupulate, inflated, sagittally compressed, (7–)8–12 mm, sparsely short villous-glandular or glabrous, throat not closing, lateral lobes shallowly convex-mucronulate, adaxial ovate with apex rounded. |
nodding, ovoid-campanulate, inflated, sagittally compressed, 9–16 mm, moderately villous-glandular, hairs gland-tipped, or mixed hirtellous and stipitate-glandular, throat closing. |
Capsules | included, (4.5–)5–8 mm. |
included, 5–12 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 30. |
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Erythranthe geyeri |
Erythranthe arenicola |
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Phenology | Flowering May–Aug(–Oct). | Flowering Apr–Aug. |
Habitat | Edges of flowing streams, marsh edges, drainage ditches, seepage areas, springs, muddy or moist banks. | Sandy beaches, especially in moist hollows among dunes, sea cliff bases, chaparral near beaches, mudstone outcrops. |
Elevation | 200–2500 m. (700–8200 ft.) | 0–100 m. (0–300 ft.) |
Distribution |
AZ; CO; IA; IL; KS; MI; MN; MO; NE; NM; OK; PA; SD; TX; WI; WY; AB; MB; ON; QC; SK; Mexico (Chihuahua, Coahuila, Distrito Federal, Durango, Hidalgo, Nuevo León, México, Querétaro, San Luis Potosí, Sonora, Veracruz, Zacatecas)
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CA |
Discussion | Erythranthe geyeri has commonly been regarded as conspecific with E. glabrata (Kunth) G. L. Nesom (as Mimulus glabratus var. jamesii), but typical E. glabrata has a different chromosome number and distinct morphology and its range does not reach the United States. In Mexico, the two species are broadly sympatric without intermediates. An allozyme study of the M. glabratus complex (R. K. Vickery 1990) indicated that the Great Plains populations of E. geyeri are distinct from those in New Mexico and Mexico, corresponding to a difference in pedicel vestiture. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
F. W. Pennell (1947, 1951) considered Erythranthe arenicola an endemic of Monterey County, but plants from adjacent San Luis Obispo and Santa Cruz counties also belong here. Most of the localities are at seaside, but some are more than a mile inland. Erythranthe arenicola is hypothesized here to be a derivative of E. guttata or E. grandis, retaining the herkogamous breeding system of its putative ancestor but reduced in size and duration. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 406. | FNA vol. 17, p. 412. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus geyeri, M. glabratus var. fremontii, M. glabratus var. jamesii, M. glabratus var. oklahomensis, M. jamesii, M. jamesii var. fremontii | Mimulus guttatus subsp. arenicola |
Name authority | (Torrey) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) |
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