Erythranthe geniculata |
Erythranthe taylorii |
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bent-stem monkeyflower, Dudley's monkeyflower |
Shasta limestone monkeyflower, Taylor's or Shasta limestone monkeyflower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Annuals, filiform-taprooted. |
Stems | ascending to decumbent or prostrate, geniculate at nodes, simple or diffusely branched, 5–60 cm, moderately villous, hairs 0.8–2 mm, multicellular, eglandular and also 0.1–0.3 mm, stipitate-glandular. |
erect, straight at nodes, simple or few-branched from base, 5–10 cm, sparsely eglandular-villous proximally, becoming sparsely short stipitate-glandular distally. |
Leaves | basal and cauline, basal usually deciduous by flowering; petiole 2–10(–35) mm; blade pinnately to subpinnately veined, broadly ovate or elliptic-ovate to triangular, 8–35 × 5–30 mm, base cuneate to rounded or subcordate, margins serrate or dentate, teeth 3–10 per side, apex acute to obtuse or rounded, surfaces moderately villous, hairs 0.8–2 mm, multicellular, eglandular, and 0.1–0.3 mm, stipitate-glandular. |
usually cauline, basal not persistent, largest at mid stem or basal and mid stem to nearly even-sized; petiole 3–5(–8) mm; blade often purple adaxially, palmately 3–5-veined, broadly ovate to elliptic-ovate or ovate-lanceolate, 4–20 × 4–12 mm, base rounded to truncate, margins serrate-dentate, teeth 2–4 per side, shallow, apex rounded to obtuse or acute, surfaces: distals moderately short-stipitate-glandular. |
Flowers | herkogamous, (1–)6–20, from all or medial to distal nodes. |
herkogamous, sometimes plesiogamous, 2–6(–8), from proximal to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, without white patches, throat red-spotted, spots concentrated or becoming coalescent into a somewhat discrete splotch at base of each of 3 abaxial lobes and sometimes 2 adaxial, bilaterally symmetric, ± bilabiate; tube-throat cylindric, 9–12 mm, exserted beyond calyx margin; limb expanded 10–18 mm diam. |
yellow, throat ceiling sometimes red-spotted or -lined, abaxial limb yellow or with 1 or 2 red splotches, bilaterally symmetric, bilabiate; tube-throat funnelform, 5–7 mm, exserted beyond calyx margin. |
Fruiting pedicels | 12–26(–55) mm, moderately villous, hairs 0.8–2 mm, multicellular, eglandular and also 0.1–0.3 mm, stipitate-glandular. |
divergent to arcuate-divergent, 6–13 mm. |
Fruiting calyces | red-spotted, campanulate-cylindric, weakly inflated, (5–)6–8 mm, margins distinctly toothed or lobed, sparsely to moderately villous-glandular, ribs shallowly wing-angled, lobes pronounced, erect to spreading or spreading-recurving. |
wing-angled, tubular-campanulate, 4–5 mm, margins distinctly toothed or lobed, densely invested with tiny, waxy-white, eglandular, papillose hairs between angles, lobes pronounced, erect. |
Capsules | included, 4–6(–7) mm. |
included, 3–4 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
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Erythranthe geniculata |
Erythranthe taylorii |
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Phenology | Flowering (Mar–)Apr–Jul. | Flowering Feb–May. |
Habitat | Granite crevices, canyon slopes, talus, crevices in volcanic outcrops, edges of boulders, roadsides, damp sandy soils, sandy water edges, gravelly soils and creek bottoms. | Crevices in limestone cliff faces and outcrops. |
Elevation | 200–900(–1200) m. (700–3000(–3900) ft.) | 900–1100 m. (3000–3600 ft.) |
Distribution |
CA
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CA
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Discussion | Erythranthe geniculata is known from an apparently disjunct cluster of populations in Butte, Sutter, and Yuba counties and then from Tuolumne and Stanislaus counties south to Kern County. Erythranthe geniculata, compared to E. floribunda, has larger, chasmogamous, and allogamous flowers. The anther pairs of E. geniculata are at different levels, and the stigma is slightly above the adaxial anther pair; in E. floribunda both anther pairs and the stigma are at the same level. Erythranthe arenaria, E. geniculata, and E. norrisii constitute a group of apparently closely related species within sect. Mimulosma endemic along the Sierra Nevada. All have ovate-petiolate leaves (only the basal ones are sometimes ovate in E. arenaria) with pinnate to subpinnate venation. The more widespread E. floribunda, which is part of the above group, also is similar, but all three endemics have larger corollas with the tube exserted at greater length beyond the calyx margin. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe taylorii is known only from the Shasta Lake region of northwestern Shasta County. Its broad, distinctly bilabiate corollas and ovate leaf blades with palmate venation are similar to those of species of the northern group of sect. Mimulosma, the "Columbia River clade" (J. B. Whittall et al. 2006) of Idaho, Montana, Oregon, Washington, and Wyoming, particularly to the Idaho endemic E. ampliata. Erythranthe taylorii is distinct from E. ampliata in its larger, papillose calyces, shorter fruiting pedicels, corollas with shorter tube-throats, and shorter capsules. Considerable corolla color variation exists in E. taylorii in the occurrence and density of red dots and lines on the throat ceiling and larger red splotches on the abaxial limb. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 404. | FNA vol. 17, p. 398. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus geniculatus, M. dudleyi, M. floribundus var. geniculatus | |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | G. L. Nesom: Phytoneuron 2013-43: 6, figs. 5–7. (2013) — (as taylori) |
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