Erythranthe geniculata |
Erythranthe lewisii |
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bent-stem monkeyflower, Dudley's monkeyflower |
great purple monkey-flower, great purple or Lewis' monkeyflower, Lewis' monkey flower, pink monkey-flower, purple monkey-flower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Perennials, rhizomatous. |
Stems | ascending to decumbent or prostrate, geniculate at nodes, simple or diffusely branched, 5–60 cm, moderately villous, hairs 0.8–2 mm, multicellular, eglandular and also 0.1–0.3 mm, stipitate-glandular. |
erect, usually simple, (15–)25–60(–75) cm, stipitate-glandular to glandular-villous. |
Leaves | basal and cauline, basal usually deciduous by flowering; petiole 2–10(–35) mm; blade pinnately to subpinnately veined, broadly ovate or elliptic-ovate to triangular, 8–35 × 5–30 mm, base cuneate to rounded or subcordate, margins serrate or dentate, teeth 3–10 per side, apex acute to obtuse or rounded, surfaces moderately villous, hairs 0.8–2 mm, multicellular, eglandular, and 0.1–0.3 mm, stipitate-glandular. |
cauline; petiole 0 mm; blade palmately veined, elliptic to ovate, ovate-lanceolate, or broadly oblanceolate, (10–)25–75(–90) × 5–35 mm, base rounded to cuneate, subclasping, margins denticulate, subentire, or entire, apex acute, surfaces stipitate-glandular to glandular-villous. |
Flowers | herkogamous, (1–)6–20, from all or medial to distal nodes. |
herkogamous, 2–6(–10), axillary at leafy medial to distal nodes. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, without white patches, throat red-spotted, spots concentrated or becoming coalescent into a somewhat discrete splotch at base of each of 3 abaxial lobes and sometimes 2 adaxial, bilaterally symmetric, ± bilabiate; tube-throat cylindric, 9–12 mm, exserted beyond calyx margin; limb expanded 10–18 mm diam. |
purple, rarely crimson, pale violet, white, pinkish white, yellowish white, or lavender, sometimes lined with red dots, bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 22–28 mm, exserted beyond calyx margins; lobe apex usually truncate to shallowly convex, shallowly retuse, throat open. |
Fruiting pedicels | 12–26(–55) mm, moderately villous, hairs 0.8–2 mm, multicellular, eglandular and also 0.1–0.3 mm, stipitate-glandular. |
(25–)35–70 mm, stipitate-glandular to glandular-villous. |
Fruiting calyces | red-spotted, campanulate-cylindric, weakly inflated, (5–)6–8 mm, margins distinctly toothed or lobed, sparsely to moderately villous-glandular, ribs shallowly wing-angled, lobes pronounced, erect to spreading or spreading-recurving. |
broadly cylindric-campanulate, not inflated, 15–22 mm, stipitate-glandular to glandular-villous, tube 12–15(–17) × 9–12 mm. |
Capsules | included, 4–6(–7) mm. |
included, 6–11 mm. |
Anthers | included, glabrous. |
included, white-villous, thecae spreading. |
2n | = 32. |
= 16. |
Erythranthe geniculata |
Erythranthe lewisii |
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Phenology | Flowering (Mar–)Apr–Jul. | Flowering Jun–Sep. |
Habitat | Granite crevices, canyon slopes, talus, crevices in volcanic outcrops, edges of boulders, roadsides, damp sandy soils, sandy water edges, gravelly soils and creek bottoms. | Stream banks, springs, wet meadows, subalpine slopes, talus, crevices, ditches. |
Elevation | 200–900(–1200) m. (700–3000(–3900) ft.) | 600–2900(–3200) m. (2000–9500(–10500) ft.) |
Distribution |
CA
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AK; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
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Discussion | Erythranthe geniculata is known from an apparently disjunct cluster of populations in Butte, Sutter, and Yuba counties and then from Tuolumne and Stanislaus counties south to Kern County. Erythranthe geniculata, compared to E. floribunda, has larger, chasmogamous, and allogamous flowers. The anther pairs of E. geniculata are at different levels, and the stigma is slightly above the adaxial anther pair; in E. floribunda both anther pairs and the stigma are at the same level. Erythranthe arenaria, E. geniculata, and E. norrisii constitute a group of apparently closely related species within sect. Mimulosma endemic along the Sierra Nevada. All have ovate-petiolate leaves (only the basal ones are sometimes ovate in E. arenaria) with pinnate to subpinnate venation. The more widespread E. floribunda, which is part of the above group, also is similar, but all three endemics have larger corollas with the tube exserted at greater length beyond the calyx margin. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe lewisii in California occurs in Modoc, Siskiyou, and Trinity counties. California records from Shasta County and south previously identified as E. lewisii are identified here as E. erubescens. The record for E. lewisii in Alaska is documented by this collection: Hyder [noted on handwritten label as "New to Alaska, Standley"], damp land, 27 June 1924, K. Whited 1291 (MO). Apparent exceptions to the characteristic flower color are these: white to lavender in Nevada (Clark County, Charleston Mountains, Train 2068, MO); pinkish white in Washington (Skamania and Yakima counties, Mt. Paddo, Suksdorf 5779, MO); white or tinged with yellow, in Wyoming (Teton County, as described by Nelson in the protologue of Mimulus lewisii var. tetonensis). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 404. | FNA vol. 17, p. 392. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus geniculatus, M. dudleyi, M. floribundus var. geniculatus | Mimulus lewisii, M. lewisii var. tetonensis |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | (Pursh) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012) |
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