Erythranthe geniculata |
Erythranthe corallina |
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bent-stem monkeyflower, Dudley's monkeyflower |
coralline monkeyflower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Perennials, rhizomatous, rhizomes often forming a mass, usually branching, filiform. |
Stems | ascending to decumbent or prostrate, geniculate at nodes, simple or diffusely branched, 5–60 cm, moderately villous, hairs 0.8–2 mm, multicellular, eglandular and also 0.1–0.3 mm, stipitate-glandular. |
usually erect to ascending-erect, few-branched, 6–25(–38) cm, moderately hirsute to hirtellous, hairs deflexed. |
Leaves | basal and cauline, basal usually deciduous by flowering; petiole 2–10(–35) mm; blade pinnately to subpinnately veined, broadly ovate or elliptic-ovate to triangular, 8–35 × 5–30 mm, base cuneate to rounded or subcordate, margins serrate or dentate, teeth 3–10 per side, apex acute to obtuse or rounded, surfaces moderately villous, hairs 0.8–2 mm, multicellular, eglandular, and 0.1–0.3 mm, stipitate-glandular. |
basal and cauline, becoming larger distally or even-sized; petiole 0 mm or proximals 1–15 mm; blade palmately 5-veined, ovate to broadly ovate, 15–45 mm, base mostly truncate to shallowly cordate, margins sharply dentate-serrate, apex obtuse, surfaces hirtellous to softly hirsute, hairs ascending, straight, dull gray, sharp-pointed, thick-walled, eglandular. |
Flowers | herkogamous, (1–)6–20, from all or medial to distal nodes. |
herkogamous, 1–3(–6), commonly solitary or from distal nodes. |
Styles | glabrous. |
sparsely hirtellous. |
Corollas | yellow, without white patches, throat red-spotted, spots concentrated or becoming coalescent into a somewhat discrete splotch at base of each of 3 abaxial lobes and sometimes 2 adaxial, bilaterally symmetric, ± bilabiate; tube-throat cylindric, 9–12 mm, exserted beyond calyx margin; limb expanded 10–18 mm diam. |
yellow, red-spotted, bilaterally symmetric, bilabiate; tube-throat narrowly funnelform to broadly cylindric, 13–20 mm, exserted beyond calyx margin; limb expanded 12–22 mm. |
Fruiting pedicels | 12–26(–55) mm, moderately villous, hairs 0.8–2 mm, multicellular, eglandular and also 0.1–0.3 mm, stipitate-glandular. |
(10–)25–75 mm, glabrous or puberulent proximally, hairs stipitate-glandular. |
Fruiting calyces | red-spotted, campanulate-cylindric, weakly inflated, (5–)6–8 mm, margins distinctly toothed or lobed, sparsely to moderately villous-glandular, ribs shallowly wing-angled, lobes pronounced, erect to spreading or spreading-recurving. |
sometimes purple-spotted, broadly cylindric-campanulate, inflated, sagittally compressed, 11–15 mm, glabrous, throat not closing, proximal lobe pair slightly upcurving. |
Capsules | included, 4–6(–7) mm. |
included, stipitate, 7–10 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
= 48, 56. |
Erythranthe geniculata |
Erythranthe corallina |
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Phenology | Flowering (Mar–)Apr–Jul. | Flowering (May–)Jun–Aug. |
Habitat | Granite crevices, canyon slopes, talus, crevices in volcanic outcrops, edges of boulders, roadsides, damp sandy soils, sandy water edges, gravelly soils and creek bottoms. | Creek banks, moraine water courses, bogs, marshes, wet meadows, roadside ditches. |
Elevation | 200–900(–1200) m. (700–3000(–3900) ft.) | (1400–)1700–2700(–3000) m. ((4600–)5600–8900(–9800) ft.) |
Distribution |
CA
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CA; NV |
Discussion | Erythranthe geniculata is known from an apparently disjunct cluster of populations in Butte, Sutter, and Yuba counties and then from Tuolumne and Stanislaus counties south to Kern County. Erythranthe geniculata, compared to E. floribunda, has larger, chasmogamous, and allogamous flowers. The anther pairs of E. geniculata are at different levels, and the stigma is slightly above the adaxial anther pair; in E. floribunda both anther pairs and the stigma are at the same level. Erythranthe arenaria, E. geniculata, and E. norrisii constitute a group of apparently closely related species within sect. Mimulosma endemic along the Sierra Nevada. All have ovate-petiolate leaves (only the basal ones are sometimes ovate in E. arenaria) with pinnate to subpinnate venation. The more widespread E. floribunda, which is part of the above group, also is similar, but all three endemics have larger corollas with the tube exserted at greater length beyond the calyx margin. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe corallina is a morphologically consistent entity that occurs only in the Sierra Nevada of California and adjacent Nevada (Washoe County and Carson City). Its chromosome number is reported as 2n = 48 and 56, compared to 2n = 28 and 56 in E. tilingii; identities of the E. corallina vouchers should be rechecked and additional counts made, since the occurrence of such wide dysploidy seems unlikely. Compared to the leaf blades of E. tilingii in the strict sense, those of E. corallina are relatively broader (broadly ovate to orbicular-ovate), the plants generally taller, and long-pedicellate flowers occasionally are produced from mid stem or even proximal nodes. The hirsutulous to hirsute vestiture of eglandular hairs on both leaf surfaces is a reliably diagnostic feature and usually easily observed with a 10× lens. Some plants of Erythranthe corallina from San Bernardino County, California, produce decumbent-ascending stems (4–10 cm) and ovate-triangular leaves (blade 5–10 × 3–6 mm), but the dense system of filiform rhizomes, flowers one to three, and hirtellous foliar vestiture serve to identify them. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 404. | FNA vol. 17, p. 410. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus geniculatus, M. dudleyi, M. floribundus var. geniculatus | Mimulus corallinus, M. minusculus, M. tilingii var. corallinus |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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