Erythranthe eastwoodiae |
Erythranthe scouleri |
|
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crimson monkeyflower, Eastwood's monkey-flower |
Columbia River monkey-flower, Scouler's monkey-flower |
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Habit | Perennials, stoloniferous, sometimes also rhizomatous. | Perennials, rhizomatous, producing long, sparsely leafy runners from basal nodes. |
Stems | scandent to pendent, usually simple, 5–30(–40) cm, villous-glandular to minutely stipitate-glandular, hairs often a mixture of longer and much shorter ones, gland-tipped. |
erect, simple or few-branched, 15–80 cm, glabrous. |
Leaves | cauline; petiole 0 mm; blade palmately 3-veined, flabellate distally to obovate to oblanceolate or elliptic, (5–)13–40(–55) × 8–20(–25) mm, largest near mid stem or distally, thick, base cuneate to rounded, subclasping, margins coarsely serrate on distal 1/2, apex acute, surfaces villous-glandular to minutely stipitate-glandular, hairs often a mixture of longer and much shorter ones, gland-tipped. |
cauline, basal usually not persistent; petiole: proximals and medial 10–25 mm (gradually merging into blade), distals 0 mm; blade palmately (3–)5–7-veined to subpinnate, (proximal to medial) oblong-elliptic to oblong-lanceolate, 25–60 × 8–18 mm, usually 3–4 times longer than wide, base attenuate, margins evenly, shallowly dentate or crenate to mucronate or mucronulate, teeth 10–20 per side, sometimes more deeply toothed at base, apex obtuse to acute, surfaces glabrous. |
Flowers | herkogamous, 2–8, axillary at leafy medial to distal nodes. |
herkogamous, (1 or)2–8, from distal nodes. |
Styles | glabrous. |
minutely, prominently hirsutulous to villosulous. |
Corollas | scarlet to orange-red or orange, palate red, not spotted or striped, bilaterally symmetric, strongly bilabiate; tube-throat narrowly funnelform, 20–30 mm, exserted 5–15 mm beyond calyx margin; throat open, palate puberulent. |
yellow, without red markings, bilaterally symmetric, bilabiate; tube-throat funnelform, 20–24 mm, exserted 10–15 mm beyond calyx margin; limb expanded 22–30 mm. |
Fruiting pedicels | 10–30(–40) mm. |
20–25 mm, glabrous. |
Fruiting calyces | cuneate-cylindric to cylindric, weakly or not inflated, 15–23(–27) mm, glabrous or minutely stipitate-glandular to sparsely glandular-villosulous, lobes triangular-acuminate. |
ovoid, inflated, sagittally compressed, 13–14 mm, glabrous, throat closing. |
Capsules | included, 6–10 mm. |
unknown. |
Anthers | exserted, villous, thecae reflexed 45º. |
included, glabrous. |
2n | = 16. |
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Erythranthe eastwoodiae |
Erythranthe scouleri |
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Phenology | Flowering May–Sep(–Nov). | Flowering May–Aug. |
Habitat | Seepages in sandstone overhangs, cave roofs, walls, crevices, and cliff bases, pinyon-juniper woodlands. | Salt marshes, river banks. |
Elevation | 900–2000 m. (3000–6600 ft.) | 0–20 m. (0–100 ft.) |
Distribution |
AZ; CO; NM; UT
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OR; WA |
Discussion | The range of Erythranthe eastwoodiae appears to be essentially contiguous with that of E. verbenacea in the Grand Canyon region, but there is no evidence of hybridization. Erythranthe eastwoodiae (as Mimulus eastwoodiae) is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe scouleri is distinctive in its oblong-elliptic leaves with long-tapering bases and closely toothed margins, completely glabrous vestiture, long, leafy runners from basal cauline nodes, large corollas with broad limbs, and prominently hairy styles. Several features suggest a close relationship to E. decora, particularly its very large corollas, hairy styles, closely toothed leaf margins, tall, simple, and erect stems, numerous runners, and its geographic range. All collections apparently have been made near the mouth of the Columbia River in Clatsop and Columbia counties, Oregon (G. L. Nesom 2013d). Recent observations (Alexander John Wright, pers. comm.; photos http://www.inaturalist.org/observations/3537008) indicate that it also occurs in Wahkiakum County, Washington, in the Julia Butler Hansen Refuge. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 394. | FNA vol. 17, p. 415. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus eastwoodiae | Mimulus scouleri, M. guttatus subsp. scouleri |
Name authority | (Rydberg) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012) | (Hooker) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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