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crimson monkeyflower, Eastwood's monkey-flower

mimule musqué, musk monkeyflower, musk-flower, musk-plant, sticky monkey-flower

Habit Perennials, stoloniferous, sometimes also rhizomatous. Perennials, rhizomatous, rooting at proximal nodes.
Stems

scandent to pendent, usually simple, 5–30(–40) cm, villous-glandular to minutely stipitate-glandular, hairs often a mixture of longer and much shorter ones, gland-tipped.

erect, sometimes ascending to decumbent, simple or branched, (2–)5–20 cm, nodes 2–4(or 5), glabrate to glandular-villous, hairs 0.5–2 mm, gland-tipped, internodes evident.

Leaves

cauline;

petiole 0 mm;

blade palmately 3-veined, flabellate distally to obovate to oblanceolate or elliptic, (5–)13–40(–55) × 8–20(–25) mm, largest near mid stem or distally, thick, base cuneate to rounded, subclasping, margins coarsely serrate on distal 1/2, apex acute, surfaces villous-glandular to minutely stipitate-glandular, hairs often a mixture of longer and much shorter ones, gland-tipped.

usually cauline, basal not persistent, distinctly separated;

petiole 0 mm or (0.5–)1–5(–10) mm;

blade pinnately veined, oblong-ovate to ovate, (10–)15–40(–50) × 5–25 mm, base obtuse-cuneate to truncate, rounded or subcordate, subclasping to sessile, margins coarsely serrate-dentate to denticulate or subentire, apex acute to obtuse, surfaces glabrate to glandular-villous.

Flowers

herkogamous, 2–8, axillary at leafy medial to distal nodes.

herkogamous, 1–8, from medial to distal nodes.

Styles

glabrous.

glabrous.

Corollas

scarlet to orange-red or orange, palate red, not spotted or striped, bilaterally symmetric, strongly bilabiate;

tube-throat narrowly funnelform, 20–30 mm, exserted 5–15 mm beyond calyx margin;

throat open, palate puberulent.

yellow, throat with fine red to blackish or brown lines extending onto lobes, red to brown dots in throat and lobes present or absent, bilaterally or nearly radially symmetric, bilabiate or nearly regular;

tube-throat narrowly funnelform, 11–18 mm, exserted beyond calyx margin;

lobes oblong-obovate, apex rounded to notched.

Fruiting pedicels

10–30(–40) mm.

(7–)10–25 mm, glabrate to glandular-villous.

Fruiting calyces

cuneate-cylindric to cylindric, weakly or not inflated, 15–23(–27) mm, glabrous or minutely stipitate-glandular to sparsely glandular-villosulous, lobes triangular-acuminate.

ridge- to wing-angled, campanulate to cylindric-campanulate, weakly or not inflated, 6–13 mm, villous to glandular-villous, lobes erect to spreading-recurving, strongly unequal to subequal, triangular to linear-lanceolate or narrowly triangular-acuminate, 2–4 mm, apex acute to obtuse.

Capsules

included, 6–10 mm.

included, 6–8 mm.

Anthers

exserted, villous, thecae reflexed 45º.

included, glabrous or slightly hirtellous to scabrous.

2n

= 16.

= 32.

Erythranthe eastwoodiae

Erythranthe moschata

Phenology Flowering May–Sep(–Nov). Flowering May–Aug.
Habitat Seepages in sandstone overhangs, cave roofs, walls, crevices, and cliff bases, pinyon-juniper woodlands. Springs and seeps, creek edges, moist meadows, ditches, along trails, roadsides, rocky ridges, granite outcrops, shaded and wet places in sagebrush, aspen, fir, spruce-fir, lodgepole pine forests, meadows.
Elevation 900–2000 m. (3000–6600 ft.) (300–)400–3100 m. ((1000–)1300–10200 ft.)
Distribution
from FNA
AZ; CO; NM; UT
[WildflowerSearch map]
from FNA
CA; CO; CT; ID; MA; ME; MI; MT; NH; NJ; NV; NY; OR; PA; RI; UT; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM [Introduced in South America (Chile), Europe, e Asia (Japan), Pacific Islands (New Zealand), Australia]
[WildflowerSearch map]
Discussion

The range of Erythranthe eastwoodiae appears to be essentially contiguous with that of E. verbenacea in the Grand Canyon region, but there is no evidence of hybridization.

Erythranthe eastwoodiae (as Mimulus eastwoodiae) is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Earlier segregation of Erythranthe moniliformis as distinct from E. moschata (for example, G. L. Nesom 2012g) emphasized a primarily erect habit and tendency toward sessile to subsessile and more densely arranged cauline leaves in E. moniliformis versus a decumbent to procumbent habit and consistently petiolate leaves on longer internodes in E. moschata. Discontinuities in morphology, geography, and ecology were not confirmed in later study by Nesom (2017). Rhizomes with small, tuberlike swellings can be observed over the whole moschata/moniliformis range, and there apparently are no consistent distinctions in vestiture and corolla size.

Mimulus acutidens Reiche (1911), a later homonym of M. acutidens Greene, pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 394. FNA vol. 17, p. 401.
Parent taxa Phrymaceae > Erythranthe Phrymaceae > Erythranthe
Sibling taxa
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms Mimulus eastwoodiae Mimulus moschatus, E. inodora, E. moniliformis, M. crinitus, M. guttatus var. moschatus, M. inodorus, M. leibergii, M. macranthus, M. moniliformis, M. moschatus var. longiflorus, M. moschatus var. moniliformis, M. moschatus var. pallidiflorus
Name authority (Rydberg) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012) (Douglas ex Lindley) G. L. Nesom: Phytoneuron 2012-39: 38. (2012)
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