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crimson monkeyflower, Eastwood's monkey-flower

disappearing monkey-flower, ephemeral monkeyflower

Habit Perennials, stoloniferous, sometimes also rhizomatous. Annuals, fibrous-rooted or filiform-taprooted.
Stems

scandent to pendent, usually simple, 5–30(–40) cm, villous-glandular to minutely stipitate-glandular, hairs often a mixture of longer and much shorter ones, gland-tipped.

erect to ascending, straight or geniculate at nodes, simple or branched at proximal and medial nodes, 6–20(–25) cm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped.

Leaves

cauline;

petiole 0 mm;

blade palmately 3-veined, flabellate distally to obovate to oblanceolate or elliptic, (5–)13–40(–55) × 8–20(–25) mm, largest near mid stem or distally, thick, base cuneate to rounded, subclasping, margins coarsely serrate on distal 1/2, apex acute, surfaces villous-glandular to minutely stipitate-glandular, hairs often a mixture of longer and much shorter ones, gland-tipped.

usually cauline, basal usually deciduous by flowering;

petiole: proximals 1–3 mm, distals 0 mm;

blade palmately 3–5-veined, narrowly ovate or narrowly lanceolate to elliptic or elliptic-lanceolate, largest 8–18(–30) × (1–)3–7 mm, relatively even-sized, or slightly reduced distally, base attenuate to obtuse or rounded, margins entire, mucronulate, or denticulate, apex acute to obtuse, surfaces minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped.

Flowers

herkogamous, 2–8, axillary at leafy medial to distal nodes.

plesiogamous, 10–20, from medial to distal nodes.

Styles

glabrous.

glabrous.

Corollas

scarlet to orange-red or orange, palate red, not spotted or striped, bilaterally symmetric, strongly bilabiate;

tube-throat narrowly funnelform, 20–30 mm, exserted 5–15 mm beyond calyx margin;

throat open, palate puberulent.

yellow to pale yellow, sparsely red-spotted or not, bilaterally symmetric, weakly bilabiate;

tube-throat cylindric, 5–8 mm, exserted 1–3 mm beyond calyx margin;

limb barely widened, lobes broadly obovate, apex rounded or mucronate.

Fruiting pedicels

10–30(–40) mm.

straight, 7–18 mm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped.

Fruiting calyces

cuneate-cylindric to cylindric, weakly or not inflated, 15–23(–27) mm, glabrous or minutely stipitate-glandular to sparsely glandular-villosulous, lobes triangular-acuminate.

winged, plicate-angled, maturing ovoid-ellipsoid to campanulate or broadly urceolate, distinctly inflated, 7–11 mm, margins distinctly toothed or lobed, sparsely, minutely hirtellous, eglandular, lobes pronounced, erect.

Capsules

included, 6–10 mm.

included, 5–9 mm.

Anthers

exserted, villous, thecae reflexed 45º.

included, glabrous.

2n

= 16.

Erythranthe eastwoodiae

Erythranthe inflatula

Phenology Flowering May–Sep(–Nov). Flowering Jun–Jul.
Habitat Seepages in sandstone overhangs, cave roofs, walls, crevices, and cliff bases, pinyon-juniper woodlands. Drying edges, banks, and beds of summer-dry watercourses, near drying edges of small lakes or impoundments, often among rocks and shoreline detritus, occasionally in moist protected areas beneath low shrubs.
Elevation 900–2000 m. (3000–6600 ft.) 1200–1700 m. (3900–5600 ft.)
Distribution
from FNA
AZ; CO; NM; UT
[WildflowerSearch map]
from FNA
CA; ID; NV; OR
Discussion

The range of Erythranthe eastwoodiae appears to be essentially contiguous with that of E. verbenacea in the Grand Canyon region, but there is no evidence of hybridization.

Erythranthe eastwoodiae (as Mimulus eastwoodiae) is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

No natural occurrences of Erythranthe inflatula are known from Washington; the type collection from Klickitat County is from a cultivated plant.

Morphological and molecular data (R. J. Meinke 1995; P. M. Beardsley et al. 2004) indicate that Erythranthe inflatula originated as a hybrid between E. breviflora and E. latidens. Its geography and biology suggest that it is reproductively stable. The putative parents are geographically and ecologically separated for most of their ranges, and the range of E. inflatula is considerably broader than the relatively small region where the parents are sympatric. In the region of sympatry, however, E. inflatula may be difficult to distinguish from one or both of its putative parents. G. L. Nesom (2012g) was not able to find morphology that would distinguish the recently described Mimulus evanescens from E. inflatula.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 394. FNA vol. 17, p. 400.
Parent taxa Phrymaceae > Erythranthe Phrymaceae > Erythranthe
Sibling taxa
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms Mimulus eastwoodiae Mimulus inflatulus, M. evanescens
Name authority (Rydberg) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012) (Suksdorf) G. L. Nesom: Phytoneuron 2012-39: 38. (2012)
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