Erythranthe eastwoodiae |
Erythranthe inamoena |
|
---|---|---|
crimson monkeyflower, Eastwood's monkey-flower |
Texas monkeyflower |
|
Habit | Perennials, stoloniferous, sometimes also rhizomatous. | Annuals, fibrous-rooted, rooting at proximal nodes, sometimes forming matlike colonies. |
Stems | scandent to pendent, usually simple, 5–30(–40) cm, villous-glandular to minutely stipitate-glandular, hairs often a mixture of longer and much shorter ones, gland-tipped. |
decumbent to decumbent-ascending basally, becoming fully erect at least in inflorescence, simple, sometimes few-branched from proximal nodes, usually distinctly fistulose, 10–30 cm, glabrous. |
Leaves | cauline; petiole 0 mm; blade palmately 3-veined, flabellate distally to obovate to oblanceolate or elliptic, (5–)13–40(–55) × 8–20(–25) mm, largest near mid stem or distally, thick, base cuneate to rounded, subclasping, margins coarsely serrate on distal 1/2, apex acute, surfaces villous-glandular to minutely stipitate-glandular, hairs often a mixture of longer and much shorter ones, gland-tipped. |
basal and cauline, basal sometimes deciduous by flowering; petiole: basal and proximals to mid cauline 10–70 mm, distals 0 mm; blade palmately veined, ovate to broadly ovate or elliptic-ovate, becoming subreniform distally, (5–)15–35(–60) mm, distal closely paired, auriculate-subclasping, base truncate to subcordate, margins dentate-serrate to shallowly dentate, teeth 5–11 per side, apex obtuse to rounded, surfaces glabrous. |
Flowers | herkogamous, 2–8, axillary at leafy medial to distal nodes. |
plesiogamous, (6–)8–18(–24), often produced from all nodes, loosely to densely racemose, chasmogamous. |
Styles | glabrous. |
glabrous. |
Corollas | scarlet to orange-red or orange, palate red, not spotted or striped, bilaterally symmetric, strongly bilabiate; tube-throat narrowly funnelform, 20–30 mm, exserted 5–15 mm beyond calyx margin; throat open, palate puberulent. |
yellow, red-spotted, bilaterally symmetric, bilabiate; tube-throat cylindric, 7–11 mm, exserted (1–)2–3(–4) mm beyond calyx margin; limb slightly expanded. |
Fruiting pedicels | 10–30(–40) mm. |
9–20 mm, glabrous, rarely sparsely stipitate-glandular. |
Fruiting calyces | cuneate-cylindric to cylindric, weakly or not inflated, 15–23(–27) mm, glabrous or minutely stipitate-glandular to sparsely glandular-villosulous, lobes triangular-acuminate. |
purple-spotted, sometimes greenish, broadly cylindric-campanulate, inflated, sagittally compressed, 7–11 mm, glabrous, rarely minutely scabrous-hirtellous or sparsely stipitate-glandular, throat not closing, abaxial lobe slightly upcurving 10–45º, spreading 45º, or sometimes deflexed 40º. |
Capsules | included, 6–10 mm. |
included, 4.5–6 mm. |
Anthers | exserted, villous, thecae reflexed 45º. |
included, glabrous. |
2n | = 16. |
= 60. |
Erythranthe eastwoodiae |
Erythranthe inamoena |
|
Phenology | Flowering May–Sep(–Nov). | Flowering Jan–Apr(–Sep). |
Habitat | Seepages in sandstone overhangs, cave roofs, walls, crevices, and cliff bases, pinyon-juniper woodlands. | Edge of seeps and creeks, mud or gravel, shallow running water, wet crevices, canyon drainages. |
Elevation | 900–2000 m. (3000–6600 ft.) | 100–2400 m. (300–7900 ft.) |
Distribution |
AZ; CO; NM; UT
|
TX; Mexico (Chihuahua, Coahuila) |
Discussion | The range of Erythranthe eastwoodiae appears to be essentially contiguous with that of E. verbenacea in the Grand Canyon region, but there is no evidence of hybridization. Erythranthe eastwoodiae (as Mimulus eastwoodiae) is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe inamoena is distinctive in its completely glabrous herbage, small corollas, flowers in racemes mostly at distal nodes and with reduced bracts, short and open-throated fruiting calyces, erect and fistulose stems, and apparent annual duration (fibrous-rooted but usually rooting at proximal cauline nodes). Presumably because of its autogamous reproduction, Erythranthe inamoena has been confused with E. cordata, especially in the trans-Pecos region of Texas, where the two are sympatric. In contrast to E. cordata, E. inamoena usually has glabrous pedicels and calyces, flowers often produced from all nodes, smaller calyces [(7–)8–11 mm] that do not close at maturity, and corollas with a shorter tube-throat (7–11 mm). Some populations in Brewster, Presidio, and Val Verde counties are identified here as Erythranthe inamoena (based on proximal-to-distal distribution of flowers and the short mature calyces with open throat) but have sparsely stipitate-glandular pedicels and calyces. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 394. | FNA vol. 17, p. 407. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus eastwoodiae | Mimulus inamoenus, M. jamesii var. texensis |
Name authority | (Rydberg) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
Web links |