Erythranthe decora |
Erythranthe inflatula |
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mannered monkeyflower, northwestern monkey -flower, sharp-leaf monkey-flower, showy monkey-flower |
disappearing monkey-flower, ephemeral monkeyflower |
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Habit | Perennials, rhizomatous, sometimes producing numerous, long, remotely leafy runners from basal cauline nodes. | Annuals, fibrous-rooted or filiform-taprooted. |
Stems | erect, simple, 20–100 cm, distals densely, minutely hirtellous. |
erect to ascending, straight or geniculate at nodes, simple or branched at proximal and medial nodes, 6–20(–25) cm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
Leaves | cauline, basal usually not persistent; petiole: proximals 8–25 mm, midcauline 3–5 mm, distalmost 0 mm; blade palmately (3–)5–7-veined, broadly ovate-triangular to ovate-lanceolate, 20–50(–60) × 10–30(–40) mm, usually 1–2 times longer than wide, base rounded to truncate to shallowly cuneate, margins dentate, teeth sharp-pointed, apex acute, surfaces sometimes glabrate, distals densely, minutely hirtellous. |
usually cauline, basal usually deciduous by flowering; petiole: proximals 1–3 mm, distals 0 mm; blade palmately 3–5-veined, narrowly ovate or narrowly lanceolate to elliptic or elliptic-lanceolate, largest 8–18(–30) × (1–)3–7 mm, relatively even-sized, or slightly reduced distally, base attenuate to obtuse or rounded, margins entire, mucronulate, or denticulate, apex acute to obtuse, surfaces minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
Flowers | herkogamous, (1 or)2–7(–14), from distal nodes. |
plesiogamous, 10–20, from medial to distal nodes. |
Styles | minutely, prominently hirsutulous to villosulous. |
glabrous. |
Corollas | yellow to chrome yellow, throat floor and tube usually red-spotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, 18–26 mm, exserted (8–)10–15 mm beyond calyx margin; limb expanded 22–30 mm. |
yellow to pale yellow, sparsely red-spotted or not, bilaterally symmetric, weakly bilabiate; tube-throat cylindric, 5–8 mm, exserted 1–3 mm beyond calyx margin; limb barely widened, lobes broadly obovate, apex rounded or mucronate. |
Fruiting pedicels | 18–35(–40) mm, distals densely, minutely hirtellous, hairs often slightly deflexed. |
straight, 7–18 mm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped. |
Fruiting calyces | green, sometimes red-spotted, ovoid, inflated, sagittally compressed, 15–19 mm, distals densely, minutely hirtellous, throat closing, lobe margins (sinuses) villous. |
winged, plicate-angled, maturing ovoid-ellipsoid to campanulate or broadly urceolate, distinctly inflated, 7–11 mm, margins distinctly toothed or lobed, sparsely, minutely hirtellous, eglandular, lobes pronounced, erect. |
Capsules | included, 8–10 mm. |
included, 5–9 mm. |
Anthers | included, glabrous. |
included, glabrous. |
Erythranthe decora |
Erythranthe inflatula |
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Phenology | Flowering May–Aug. | Flowering Jun–Jul. |
Habitat | River banks, streamsides, dripping banks, moist meadows. | Drying edges, banks, and beds of summer-dry watercourses, near drying edges of small lakes or impoundments, often among rocks and shoreline detritus, occasionally in moist protected areas beneath low shrubs. |
Elevation | 1000–1600 m. (3300–5200 ft.) | 1200–1700 m. (3900–5600 ft.) |
Distribution |
ID; OR; WA; BC
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CA; ID; NV; OR |
Discussion | Erythranthe decora is distinct in its uniformly ovate to ovate-lanceolate leaf blades with truncate bases and regularly toothed margins, relatively long internodes, rhizomatous habit, mostly unbranched stems, often with leafy runners from basal nodes, large corollas, hairy styles, and minutely hirtellous stems, pedicels, calyces, and leaf surfaces. The densely produced rhizomes suggest a relationship with the E. tilingii group. A population in Clearwater County, Idaho, appears to be long-disjunct from the main range of the species in northwestern Oregon and the western half of Washington. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
No natural occurrences of Erythranthe inflatula are known from Washington; the type collection from Klickitat County is from a cultivated plant. Morphological and molecular data (R. J. Meinke 1995; P. M. Beardsley et al. 2004) indicate that Erythranthe inflatula originated as a hybrid between E. breviflora and E. latidens. Its geography and biology suggest that it is reproductively stable. The putative parents are geographically and ecologically separated for most of their ranges, and the range of E. inflatula is considerably broader than the relatively small region where the parents are sympatric. In the region of sympatry, however, E. inflatula may be difficult to distinguish from one or both of its putative parents. G. L. Nesom (2012g) was not able to find morphology that would distinguish the recently described Mimulus evanescens from E. inflatula. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 414. | FNA vol. 17, p. 400. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus guttatus var. decorus, M. decorus | Mimulus inflatulus, M. evanescens |
Name authority | (A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) | (Suksdorf) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) |
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